Plant Protection News Supplements N9, 2013
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- 2013-03-11
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- Diptera, Dolichopodidae, Asyndetus, Syntormon, Palaearctic, Russia, Greece, Israel, Afrotropical, D.R. Congo, Gambia, Madagascar, Namibia, Senegal, Oriental, India, new species, new records, key, check-list
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- Igor Ya. Grichanov
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Grichanov I.Ya. 2013. Systematic notes on West-Palearctic species of the genus Syntormon Loew (Diptera: Dolichopodidae). In: Grichanov I.Ya., Negrobov O.P. (Editors). Fauna and taxonomy of Dolichopodidae (Diptera). Collection of papers. St.Petersburg: VIZR RAAS (Plant Protection News Suppl., ISSN 1815-3682): 3-26.
Grichanov I.Ya. 2013. Afrotropical species of the genus Asyndetus Loew (Diptera: Dolichopodidae) with notes on some Palaearctic and Oriental species. In: Grichanov I.Ya., Negrobov O.P. (Editors). Fauna and taxonomy of Dolichopodidae (Diptera). Collection of papers. St.Petersburg: VIZR RAAS (Plant Protection News Suppl., ISSN 1815-3682): 27-46.
Negrobov O.P., Selivanova O.V., Maslova O.O., Chursina M.A. . 2013. Check-list of predatory flies of the family Dolichopodidae (Diptera) in the fauna of Russia. In: Grichanov I.Ya., Negrobov O.P. (Editors). Fauna and taxonomy of Dolichopodidae (Diptera). Collection of papers. St.Petersburg: VIZR RAAS (Plant Protection News Suppl., ISSN 1815-3682): 47-93.
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- <?xml version="1.0" encoding="UTF-8" ?> <?xpacket begin='' id='W5M0MpCehiHzreSzNTczkc9d'?> <?xpacket begin="" id="W5M0MpCehiHzreSzNTczkc9d"?> 2013-03-14T13:48:02+04:00 Acrobat PDFMaker 8.1 for Word 2013-06-04T12:41:52+04:00 2013-06-04T12:41:52+04:00 Acrobat Distiller 8.1.0 (Windows) Diptera; Dolichopodidae; Asyndetus; Syntormon; Palaearctic; Russia; Greece; Israel; Afrotropical; D.R. Congo; Gambia; Madagascar; Namibia; Senegal; Oriental; India; new species; new records; key; check-list xml I.Ya. Grichanov O.P. Negrobov M.A. Chursina O.O. Maslova O.V. Selivanova Fauna and taxonomy of Dolichopodidae (Diptera) Plant Protection News Supplement, N 9
I.Ya. Grichanov, O.P. Negrobov, M.A. Chursina, O.O. Maslova, O.V. Selivanova Diptera Dolichopodidae Asyndetus Syntormon Palaearctic Russia Greece Israel Afrotropical D.R. Congo Gambia Madagascar Namibia Senegal Oriental India new species new records key check-list uuid:9e574cbf-b723-445e-97bd-111317ec9b6d uuid:da3dc5b0-0548-4164-a07d-c5e8c5dea260 1 B True http://www.vestnik.iczr.ru/supplement/index.html Print <?xpacket end="w"?> <?xpacket end='r'?>
All-Russian Institute of Plant Protection RAAS
ISSN 1815-3682
ÂÅÑÒÍÈÊ ÇÀÙÈÒÛ ÐÀÑÒÅÍÈÉ
Приложение
Supplement
Fauna and taxonomy of Dolichopodidae (Diptera) Collection of papers St.Petersburg 2013Фауна и таксономия хищных мух Dolichopodidae (Diptera). Сборник научных работ. Под редакцией И.Я. Гричанова и О.П. Негробова. Санкт-Петербург: ВИЗР РАСХН, 2013. 96
c. (Приложение к журналу «Вестник защиты растений»).
Fauna and taxonomy of Dolichopodidae (Diptera). Collection of papers. Igor Ya. Grichanov & Oleg P. Negrobov, editors. St.Petersburg: VIZR RAAS, 2013. 96 p. («Plant Protection News, Supplement»).
Сборник включает обзорные статьи по фауне и таксономии хищных мух-зеленушек семейства Dolichopodidae. Описаны новые виды, приведены новые указания для видов из Палеарктической, Ориентальной и Афротропической зоогеографических областей. Составлены региональные определители видов из родов Asyndetus и Syntormon. Впервые составлен справочный список 52 родов, 735 видов и подвидов семейства Dolichopodidae, отмеченных на территории Российской Федерации. Сборник будет полезен специалистам – энтомологам и экологам, интересующимся энтомофагами, студентам и аспирантам учебных и научных учреждений.
Работа выполнена в рамках Программы фундаментальных научных исследований в Российской Федерации на долгосрочный период (2013– 2020 годы) (утверждена распоряжением Правительства Российской Федерации от 27 декабря 2012 г. № 2538-р). Издание поддержано грантом РФФИ (11-04-01051а).
Рекомендовано к печати Ученым советом Всероссийского научноисследовательского института защиты растений РАСХН 11 марта 2013 г.
Copyright 2013© I.Ya. Grichanov, O.P. Negrobov, M.A. Chursina, O.O. Maslova, O.V. Selivanova. ISSN 1815-3682
SYSTEMATIC NOTES ON WEST-PALAEARCTIC SPECIES OF THE GENUS SYNTORMON LOEW (DIPTERA: DOLICHOPODIDAE)
Igor Ya. Grichanov
All-Russian Institute of Plant Protection (VIZR), Podbelskogo 3, St.Petersburg, Pushkin, 196608, Russia. E-mail: grichanov@mail.ru
Systematic information on the West-Palaearctic species of the sympycnine genus Syntormon Loew, 1857 is reviewed. A new species Syntormon pilitibia sp. n. from Israel and a new subspecies S. macula mediterraneus ssp. n. from Greece and Israel are described. The taxonomic status and characters of some species of the genus are discussed. Lectotype and paralectotypes are designated for S. cilitibia Stackelberg, 1947. A check list of West-Palaearctic species of Syntormon is compiled, as well as a revised key to 21 species and 5 firstly defined species groups (combining 18 poorly distinguishable species) based mainly on male secondary sexual characters.
KEY WORDS: Dolichopodidae, Syntormon, Palaearctic, Greece, Israel, new species, new records, key.
И.Я. Гричанов. Заметки по систематике западно-палеарктических видов рода Syntormon Loew (Diptera: Dolichopodidae)
Обобщены данные по систематике западно-палеарктических видов рода Syntormon Loew, 1857 из подсемейства мух-зеленушек Sympycninae. Описаны новый вид Syntormon pilitibia sp. n. из Израиля и новый подвид S. macula mediterraneus ssp. n. из Греции и Израиля. Обозначены лектотип и паралектотипы для S. cilitibia Stackelberg, 1947. Составлен каталог описанных западно-палеарктических видов и определитель 21 вида и 5 групп видов (включающих 18 слабо различимых видов), основанный главным образом на вторичнополовыхпризнаках самцов.
Всероссийский научно-исследовательский институт защиты растений (ВИЗР), шоссе Подбельского, 3, Санкт-Петербург, Пушкин, 196608, Россия.
Introduction
Syntormon Loew, 1857 is an unrevised genus of the subfamily Sympycninae and includes about 110 species, of which more than 50 were described in the Palaearctic Region. An outdated key of Becker (1918) comprised 16 Palaearctic species, and Parent (1938) diagnosed in his key 21 mainly European species. Negrobov (1975) keyed males of 35 Palaearctic species, of which I recognize 29 West-Palaearctic species inhabiting Europe, North Africa, West and Central Asia.
Later a number of new species were described from the Palaearctic (Vaillant, 1983; Negrobov & Shamshev, 1984, 1985; Pârvu, 1985, 1989; Negrobov & Zhilina, 1986; Yang, 1998, 1999; Yang & Saigusa, 2000; Negrobov & Kechev, 2012), and Wang et al. (2008) published a key to males of Chinese species. Speight et al. (1995) synonymised Bathycranium Strobl, 1892 and Syntormon.
The West-Palaearctic species of Syntormon now cannot be distinguished reliably based on the key published by Negrobov (1975), which was based largely on the variable characters and outdated nomenclature, with several missing old species. Here I recognize 41 (including new) West-Palaearctic species of the genus, though some of them may be placed in synonymy after a future revisional work. Two species are known from females only.
In this paper a new species Syntormon pilitibia sp. n. from Israel and a new subspecies S. macula mediterraneus ssp. n. from Greece and Israel are described. The taxonomic status and characters of some species of the genus are discussed. A check list of
4
West-Palaearctic species of Syntormon is compiled. Below I give also a revised key to 21 species and 5 firstly defined species groups (combining 18 poorly distinguishable species) based mainly on male secondary sexual characters.
Material and methods
The holotypes and paratypes of the new species and new subspecies and other material cited are housed at the Zoological Museum of Moscow State University, Russia (MZUM), at the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia (ZIN) and at the Department of Zoology, Tel Aviv University, Israel (TAU).
Specimens were studied and illustrated with a ZEISS Discovery V–12 stereomicroscope and an AxioCam MRc5 camera. Morphological terminology and abbreviations follow Grichanov (2007) and Cumming & Wood (2009). The relative lengths of the podomeres should be regarded as representative ratios and not measurements. Body length is measured from the base of the antenna to the tip of abdominal segment 7. Wing length is measured from the base to the wing apex. Male genitalia were macerated in 10% KOH. Figures showing the male genitalia in lateral view are oriented as they appear on the intact specimen, with the morphologically ventral surface of the genitalia facing up, dorsal surface down, anterior end facing right and posterior end facing left. Photos were made by the author of this paper. Information on world distribution for each species listed follows Grichanov (2003–2013).
Systematics
Family Dolichopodidae Subfamily Sympycninae Genus Syntormon Loew
Syntormon Loew, 1857: 35; Negrobov, 1975: 652; Speight et al., 1995: 356; Grichanov et al., 2011: 35. Type species: Rhaphium metathesis Loew, 1850 (designation by Coquillett, 1910: 611)
Plectropus Haliday, 1832: 353 (nec Kirby, 1826). Type species: Musca pallipes Fabricius, 1794 (designation by Westwood, 1840: 134).
Synarthrus Loew, 1857: 35; Schiner, 1862: 192. Type species: Musca pallipes Fabricius, 1794 (monotypy).
Eutarsus Loew, 1857: 45 (nec Hessling, 1852); Dyte, 1969: 46 (as synonym of subgenus Drymonoeca Becker); Negrobov, 1975: 653. Type species: Porphyrops aulica Meigen, 1824 (monotypy).
Bathycranium Strobl, 1892: 103; Speight et al., 1995: 351. Type species: Dolichopus bicolorellus Zetterstedt, 1843 (monotypy).
Drymonoeca Becker, 1907: 108; Becker, 1922: 155 (as synonym of Eutarsus Loew). Type species: Drymonoeca calcarata Becker, 1907 [=Syntormon aulicus (Meigen, 1824)] (monotypy).
Diagnosis. Usually small species; antennal scape with or without dorsal setation; pedicel with a finger-like apical process projected into basal inner concavity of post-pedicel; male postpedicel distinctly elongated, rarely short (short in females); arista-like stylus apical or subapical (sometimes dorsal in females); male tarsi often modified and/or ornamented; metepimeron more or less haired (in contrast to bare metepimeron in closely related Nearctic Parasyntormon Wheeler, 1899 and West-Palaearctic Ceratopos Vaillant, 1952).
Remarks. Proposing a new genus, Loew (1857: 35) definitely changed the name Rhaphium tarsatum (gender neuter) into Syntormon tarsatus (gender masculine), using subsequently the latter gender (e.g., Loew, 1869, 1873). Later many scientists followed the masculine gender of the genus; nevertheless, some others used the neuter gender for the Syntormon species, sometimes explaining their position (e.g., Chandler, 1998). As a result, many national list compilers were recently confused, inserting species of the genus in their lists with both neuter and masculine gender endings. I follow here original (Loew’s) proposal, using the masculine gender for all species names.
Review of West-Palaearctic species of the genus Syntormon
Syntormon abbreviatus Becker, 1918: 272. Type locality: Tunisia: Bel-Mehtia, Aegn- Draham. Distribution: Hungary, Tunisia, Turkey. Remarks: Excluded from Afrotropical Region by Grichanov (2001: 183). Here excluded from Canary Is. because S. brevicornis is here restored from synonymy to species status (see below). A record from Hungary needs confirmation.
Syntormon aulicus (Meigen), 1824: 48 (Porphyrops); Meigen, 1838: 154 (Argyra); Zetterstedt, 1843: 615 (Dolichopus); Loew, 1857: 45 (Eutarsus); Negrobov, 1975: 654 (Syntormon). Type locality: not given. Distribution: Algeria, Austria, Azerbaijan, Belgium, Bulgaria, Czech, Denmark, France, Germany, Greece, Ireland, Italy, Morocco, Poland, Russia (Moscow), Spain, Sweden, Tunisia, Turkey, UK, “Middle Asia”. =calcaratus (Becker), 1907: 109 (Drymonoeca). Type locality: Tunisia: Ain-Dram, Zaghouan. Material: 1♂, [France]: Cherbourg / Eutarsus aulicus Meig., det. O. Parent [ZIN].
Syntormon bicolorellus (Zetterstedt), 1843: 617 (Dolichopus); Haliday: in Walker et al., 1851: 212 (Porphyrops); Strobl, 1892: 103 (Bathycranium); Kowarz, 1889: 175 (Sympycnus); Schiner, 1862: 184 (Xanthochlorus); Speight et al., 1995: 356 (Syntormon); Grichanov, 2006: 183. Type locality: «Suecia meridionali et media, Scania ad Esperod, Gottlandia ad Martebo, Dania» (Sweden, Denmark). Distribution: Austria, Belgium, Czech, Denmark, Estonia, Finland, France, Germany, Hungary, Ireland, Latvia, ?Mongolia (female only), Netherlands, Norway, Poland, Romania, Russia (Leningrad, Lipetsk, Moscow, Vologda, Voronezh), Slovakia, Sweden, UK. =bicolor (Zetterstedt), 1843: 617 (Argyra) [«Staeger in litt.»]. Material: 1♂, Sweden: Uppsala, 11.VII.2002, Grichanov [ZIN]; 1♂, [France]: Aire, VI.1921 / Bathycranium bicolorellum Zett., det. O. Parent [ZIN]; 4♀, [Russia]: Yashchera, Luzhskii Distr., Leningrad Region, 21.VIII, 2.IX.1966, 14, 31.VIII.1967, Stackelberg [ZIN]; 1♂,1 ♀, Finland: Helsinki env., lake shore, 5.IX.2003, Grichanov [ZIN].
Syntormon brevicornis Frey, 1936: 67, nom. ressur.; Negrobov, 1975: 654; Negrobov, 1991: 53 (as synonym of S. abbreviatus); here restored to species status. Type locality: «Canaria: Tafira, Tümpel; Palma: La Caldera». Distribution: Spain (Canary Is.) Remarks: Negrobov (1991) placed the species in synonymy to S. abbreviatus without explanation. Negrobov (1975) in his key placed S. brevicornis near S. rufipes (=S.
pumilus). However, Frey (1936) compared his new species with S. denticulatus, distinguishing S. brevicornis (body length is 2-3 mm) in shorter postpedicel (as long as high), shorter and stronger curved hooks on posterior basitarsus. Therefore, I restore S. brevicornis from synonymy.
Syntormon bulgariensis Negrobov et Kechev, 2012: 93. Type locality: Bulgaria: Pamporovo, Rhodopes Mts. Distribution: Bulgaria.
Syntormon cilitibia Stackelberg, 1947: 96, 100. Type locality: [Tajikistan]: Khorog on Gunt River, Shugnan (originally published as Hissar range, Kondara; Varzob valley near Stalinabad; Chorog, W. Pamirs). Distribution: Afghanistan, Kyrgyzstan, Tajikistan, Uzbekistan. Material (Fig. 1): Lectotype: ♂, [Tajikistan]: Khorog on Gunt River, Shugnan, 2000 m, 25.IX.1943, Stackelberg [ZIN]. Paralectotypes: 3♂, same locality [ZIN]. Additional material: 1♂, Tajikistan: Kondara Gorge, 1100 m, Varzob Valley, Gussakovskii, 13.IX.1939 [ZIN]; 1♂, [Uzbekistan]: Shakhimardan / 8 [yellow label; ex coll. A.P. Fedtshenko 1871; ZIN]; 1♂, [Uzbekistan]: Sangy Dzhaman / 5 [orange label; ex coll.
A.P. Fedtshenko 1871; ZIN]. Remarks: Lectotype and paralectotypes are here designated to fix the current taxonomic concept and ensure consistent future interpretation. They fully correspond to the original description. Stackelberg (1947: 96) listed 8 males under S. cilitibia description, including 3 males collected in the Varzob Valley (on 18.IX.1938 and 9.X.1943) and 5 males collected at Khorog (on 25.IX.1943). A male collected on 18.IX.1938 is entirely destroyed, but bearing a white label hand-written by A.A. Stackelberg: «Syntormon cilitibia sp.n.» A male collected on 9.X.1943 was not found. The other pins had no any identification label. One male collected at Khorog is also destroyed. A.A. Stackelberg treated also A.P. Fedtshenko collection (probably before 1927) that contains (in addition to listed material from Shakhimardan and Sangy Dzhaman) three empty pins with labels «Sangy Dzhaman» and «5»; one of those pins has identification label written by A.A. Stackelberg: Syntormon [unpublished name] sp.n. Shakhimardan (or Shohimardon) is a small exclave of Uzbekistan in Fergana Province, surrounded by Kyrgyzstan. So, this is a first record of the species from Uzbekistan and Kyrgyzstan.
Syntormon codinai Parent, 1924: 9. Type locality: Morocco: Tanger, Cabo Espartel. Distribution: Morocco, Spain, Turkey. Excluded from Afrotropical Region (Grichanov, 2001: 183).
Syntormon denticulatus (Zetterstedt), 1843: 478; Loew, 1857: 34-35 (Syntormon). Type locality: Sweden: Scania. Distribution: Abkhazia, Afghanistan, Armenia, Azerbaijan, Belorussia, Bulgaria, Estonia, Finland, France, Germany, Greece, Israel, Italy, Kyrgyzstan, Norway, Poland, Romania, Russia (Adygea, Alania, Kabardino-Balkaria, Karelia, Leningrad, Moscow, Murmansk, Stavropol’), Sweden, Tajikistan, Turkey, UK, Ukraine, «North Africa». =aculeatus (Zetterstedt), 1843: 479 (Rhaphium); Becker, 1902: 54 (Syntormon); Grichanov, 2006: 183 (synonymised by Becker, 1918: 273). Type locality: Sweden: Ostrogothia, Haradshammar. =biseriatus (Loew), 1850: 123 (as a variation of Rhaphium denticulatum Zetterstedt, 1843); Loew, 1873: 249 (Syntormon) (Haliday, in Walker et al., 1851: 204
[Rhaphium]). Type locality: Germany. =pumilus Parent, 1925: 50 (nec Meigen, 1824; misident.) (Collin, 1940: 267). Material: 1♂, [France]: Cotenin, VIII.1925 / Syntormon denticulatus Zett., det. O. Parent [ZIN]; 6♂, [Tajikistan]: Stalinabad, Dyushambinka Valley, 9-25.X1943, Stackelberg [det. A. Stackelberg] [ZIN]; 1♂, [Kyrgyzstan]: Alai / 21 [yellow label; ex coll.
A.P. Fedtshenko 1871; ZIN]; 1♂, [Kyrgyzstan]: Taka [pass] / 8 [blue label; ex coll. A.P. Fedtshenko 1871; ZIN]; 2♂, 1♀, Russia: Sochi, Mt. Akhun, 22, 28.X.2006, N. Vikhrev [MZUM]. Remarks: The species was previously recorded from «Middle Asia». So, this is a first record of the species from Kyrgyzstan and Tajikistan.
Syntormon filiger Verrall, 1912: 58 (nom. nov. for Rhaphium rufipes Zetterstedt, 1849, nec Meigen, 1824, nec Zetterstedt, 1838). Type locality: Denmark: ad Hafniam, ube in Amager (automatic). Distribution: Austria, Belgium, Bulgaria, Czech, Denmark, Finland, France, Germany, Greece (North Aegean), Hungary, Kazakhstan (Astana), Russia (Astrakhan, Rostov, Novosibirsk), Netherlands, Poland, Sweden, UK, Ukraine (Crimea). =rufipes (Zetterstedt), 1849: 3060 (Rhaphium); Verrall, 1912: 58 (Syntormon). Type locality: Denmark: «ad Hafniam, ube in Amager». =obscurifrons Parent, 1932: 229 (synonymised by Parent, 1938: 453). Type locality: Austria: Burgenland, Apetlon. Material: 1♂, [Kazakhstan]: Tselinograd env., Novoishimka, 8.VI.1989, Grichanov [ZIN].
Syntormon francoisi Meuffels et Grootaert, 1999: 291 (nom. nov. for Syntormon parvus Vaillant, 1983, nec Vanschuytbroeck, 1951). Type locality: France: Alps, cascade de Ray, vallée de la Gordolasque (automatic). Distribution: France. =parvus Vaillant, 1983: 274 (nom. preoccupied, nec Vanschuytbroeck, 1951). Type locality: France: Alps, cascade de Ray, vallée de la Gordolasque. Remarks: The species may represent the true species or be placed in synonymy to either
S. fuscipes or S. spicatus as there are no clear differences between the species concepts. Describing his new species from the Maritime Alps, Vaillant (1983) compared it with a male of «S. spicatus» taken from the Romanian Carpathians, whereas the type locality of S. spicatus is probably western Poland, and that of S. fuscipes – southern Germany. See also remarks under S. spicatus.
Syntormon freymuthae Loew, 1873: 252; Becker, 1918: 276 (as a variation of S. denticulatus), rest. by Parent, 1927: 91-92. Type locality: Russia: «Nischegrod’schen Gouvernement; bei Malaja Arja in Moskauer Gouvernement». Distribution: Estonia, Russia (Leningrad, Moscow, Nizhnii Novgorod), «Middle Asia». Material: 1♂, [Russia]: Sablino, Petrograd. Gub., 9.V.1923, Stackelberg [det. A. Stackelberg] [ZIN].
Fig. 1. Syntormon cilitibia Stackelberg, habitus. Fig. 2. Syntormon pennatus Ringdahl, habitus (Kabardino-Balkaria, compared with types).
Fig. 3. Syntormon turanicus Stackelberg, habitus (holotype).
Syntormon fuscipes (von Roser), 1840: 56 (Porphyrops); Denninger, 1950: 45 (Syntormon). Type locality: not given (Germany: Württemberg). Distribution: Abkhazia, Andorra, Austria, Belgium, Bulgaria, Czech, Denmark, France, Germany, Greece, Hungary, Netherlands, Poland, Romania, Russia (Krasnodar), Slovakia, Spain, Sweden, Turkey, UK, Ukraine (Carpathians, Crimea), «Yugoslavia»; Afrotropics: Burundi, Kenya. =spicatus Becker, 1918: 283 (nec Loew, 1857; misident.). Material: 1♂, Abkhazia: Musserskii reserve, 12.VI.1978, V. Rikhter [ZIN]. Remarks: Becker (1918: 219) considered Porphyrops fuscipes as doubtful species known from Germany only. Denninger (1950) synonymised Porphyrops fuscipes and Syntormon spicatus sensu Becker, not giving redescription or figures for a type of the former species. Becker (1918: 283) gave exact illustration of what he thought S. spicatus that was considered typical by all consequent researchers. The material cited by Becker was collected from «Silesia, Hungary, Corfu and France» and may belong to different species. The origin of specimen(s) from which his figures were made is unclear. Therefore, I restore S. spicatus from synonymy and consider distributional records for this species belonging to S. fuscipes. See also remarks under S. spicatus and S. francoisi. Two phenotypes of the species were recognized in the Afrotropical Region (Grichanov, 2001). They may represent different species or subspecies, and their real degree of separation from S. fuscipes, S. francoisi or S. spicatus may require molecular investigation. This is a first record of the species from Abkhazia.
Syntormon giordanii Negrobov, in Negrobov et Matile, 1974: 842. Type locality: Iran: «Khorramshahr, Stagno Porco Com. Mar.». Distribution: Iran. Remarks: Negrobov (1991) erroneously recorded Italy as the species type locality (in fact, it is a country of types’ depository and motherland of a collector). Therefore, Grichanov (2007) excluded this species from the fauna of Europe. See also remarks under S. samarkandi.
Syntormon iranicus Negrobov, in Negrobov et Matile: 1974: 842. Type locality: Iran: «Qars-i-Shirin, 40 km Est». Distribution: Iran.
Syntormon latitarsis Negrobov et Shamshev, 1984: 49. Type locality: Russia: Caucasus, 14 km N of Krasnaya Polyana, Chugush Mt. Distribution: Russia (Krasnodar).
Syntormon luteicornis Parent, 1927: 61. Type locality: France: Apt, Vaucluse. Distribution: France. Remarks: The species was originally described by a single female from southern France. Speight et al. (1995) redescribed the holotype and doubted records of the species from other countries (Belgium, Czech, Romania, Spain). The specimen may represent an aberrant or juvenile form of a related Syntormon species with the two setae on female face
(e.g. S. tabarkae) or a female of Ceratopos seguyi Vaillant, 1952, described from northern Algeria.
Syntormon macula Parent, 1927: 57 (as macula Oldenberg). Type locality: Romania: Mehadia, Alpes de Transylvanie (Hongrie). Distribution: ?Bulgaria, Germany, Hungary, Italy, Romania, Switzerland, UK. Remarks: The species was originally described by a single female from the Transylva
nian Alps, later recorded from several more countries. Its male was included in a key to British dolichopodid flies by d’Assis Fonseca (1978), but seems to be described after single immature male only (d’Assis Fonseca, 1949). See diagnosis of S. macula mediterraneus ssp. n.
Syntormon metathesis (Loew), 1850: 118 (Rhaphium); Loew, 1857: 34-35 (Syntormon). Type locality: Germany. Distribution: Austria, Belgium, Czech, Estonia, Finland, France, Germany, Hungary, Latvia, Netherlands, Poland, Romania; Russia (Krasnodar, Leningrad, «Ural», Voronezh), Spain, Sweden, Switzerland, Slovakia, Turkey. =simplicipes Frey, 1915: 42; Becker, 1918: 279. Type locality: Finland: Karislojo, bei Pellonkyla. =dobrogicus Pârvu, 1985: 151; Grichanov, 2007: 70. Type locality: Romania: Dobrogea, Babadag. Remarks: Comparing surstylus drawn by Pârvu (1985: Fig. 3C and 1989: Fig. 4A) for S. dobrogicus, I suppose that the pictures belong to different species.
Syntormon miki Strobl, 1899: 126. Type locality: Spain: Algeciras. Distribution: Czech, France incl. Corsica, Greece incl. Crete, Ireland, ?Israel, Italy, Morocco, Poland, Portugal, Spain, «Yugoslavia», ?Slovenia, Sweden, Tunisia, UK. =florentinus Becker, 1918: 279 (as florentinus Oldenberg in litt.) (nom. nud.).
Syntormon monilis (Haliday), in Walker et al., 1851: 205 (Rhaphium); Loew, 1859: 15 (Synarthrus); Becker, 1918: 279 (Syntormon). Type locality: England; Ireland. Distribution: Algeria, Austria, Belgium, Bulgaria, Czech, Denmark, France, Germany, Hungary, Ireland, Italy, Latvia, Morocco, Netherlands, Poland, Romania; Russia (Kabardino- Balkaria, Krasnodar, Leningrad, Moscow, Ural), Slovakia, Sweden, Switzerland, Tunisia, Turkey, UK, «Yugoslavia». Material: 1♂, [Russia]: Petrograd. Gub., Kartashevka, 3.VII.1926, Stackelberg [det. A. Stackelberg] [ZIN]; 1♂, Russia: Kabardino-Balkaria, 5 km W Zalukodes, 21.VI.2000, Grichanov [ZIN].
Original description (Walker et al., 1851: 205). Male. Long. 1¼; alar. 2½ lin. Brassy-green. Front bluish. Antennae half as long as the thorax; arista nearly as long as the antennae. Wings narrowed towards the base. Abdomen with the second segment usually yellowish beneath and at the sides. Legs and fore coxae yellow; tarsi and hind femora dusky at the tip; fore metatarsus slightly dilated at the tip, last two joints of the middle tarsi dilated; hind metatarsus armed at the base beneath with a sharp hooked tooth, fringed with some short curved bristles. Rare. (E. I.).
Remarks: This short Haliday’s description was for a long time considered exact to distinguish the species from other close relatives. It is worth noting that Haliday was probably the first who noted individual variability of abdomen colour in the genus Syntormon («Abdomen with the second segment usually yellowish…»). Most of the subsequent redescriptions of S. monilis (e.g. Loew, Becker, Parent) noted entirely dark abdomen, although I keep a photo of a true Swedish S. monilis with yellow at base abdomen, and M. Pollet found the same phenotype in Belgium (see Pârvu, 2009). Unfortunately, the coloration of abdomen (and legs) was considered a key character by many Syntormon key compilers, that caused creation of some new synonymic names or names to be synonymised in the genus.
Also, the Haliday’s description did not mention the modification of the 2nd segment of fore tarsus, but describing the «fore metatarsus [=protarsus, =basitarsus, =segment 1] slightly dilated at the tip». However, in Dolichopodidae the apical dilation of one tarsomere always corresponds to basal dilation of an adjacent tarsomere. As a result, Pârvu (1989) did not take this rule into account and described S. silvianus that has no significant differences from the species concept of S. monilis except for the coloration of abdomen and hind femur. The 2nd segment of fore tarsus of S. monilis was described as globular («kugelförmig») by Loew (1859) and «swollen at base» by Parent (1938), but was considered simple (not swollen) by Pârvu (2009) who used a disputable note by Hodge (1993) on the presence of two forms of S. monilis in Britain (with or without swelling on fore tarsomere 2). Judging from a brief comparison of the two forms, I can suppose that Hodge’s ‘Species A’ is a colour variation of true S. monilis described under the name S. silvianus, whereas his ‘Species B’ belongs to fuscipes group of species with simple fore tarsus (see my remarks under S. spicatus). Pârvu (2009) erroneously regarded S. monilis to be a rare endemic of the British Isles, whereas
S. silvianus to be a widely distributed species in many countries of West Palaearctic. At present I consider the latter species as a doubtful Romanian endemic with uncertain relation to both S. monilis and Caucasian S. submonilis that was diagnosed also by variable colour characters (Negrobov, 1975: 654). Key characters of all the three species must be redescribed and accurately figured in order to define their true relations.
Syntormon mutillatus Becker, 1918: 280. Type locality: France: «Gabas, Lamus, West-Pyrenaen». Distribution: France.
Syntormon obscurior Parent, 1938: 452 (in key, as a variation of S. sulcipes) (described by Parent, 1927: 93, as unnamed variation of S. sulcipes); Negrobov, 1975: 657 (in key, as a subspecies of S. sulcipes); Negrobov, 1991: 56 (as a synonym to S. sulcipes); stat.
n. Syntypes (males and females) in M. Zerny’s collection (probably, Naturhistorisches Museum Wien, Austria). Type locality: Albania. Remarks: The name obscurior was validated by Negrobov (1975) according to ICZN,
45.6.4.1 as published before 1981. Negrobov (1991) placed the name in synonymy to S. sulcipes, supposing that the variety originated from France. However, describing a new variety, Parent (1927) stated that the material was collected in Albania.
Syntormon pallipes (Fabricius), 1794: 340 (Musca); Fabricius, 1805: 266 (Dolichopus); Meigen, 1824: 55 (Porphyrops); Haliday, 1832: 353 (Plectropus); Haliday, in Walker et al., 1851: 204 (Rhaphium); Loew, 1857: 35 (Synarthrus); Schiner, 1862: 192 (Syntormon); Grichanov, 2001: 186. Type locality: Germany. Distribution: Abkhazia, Afghanistan, Algeria, Armenia, Austria, Azerbaijan, Belgium, Bulgaria, China, Czech, Denmark, Egypt, Estonia, Finland, France, Georgia, Germany, Greece incl. Crete, Hungary, Iceland, Iran, Iraq, Ireland, Israel, Italy, Jordan, Kyrgyzstan, Latvia, Morocco, Netherlands, Norway, Poland, Portugal incl. Madeira, Azores, Romania, Russia (Adygea, Alania, Kabardino-Balkaria, Karachai-Cherkessia, Krasnodar, Leningrad, Murmansk, Rostov, Voronezh), Slovakia, ?Slovenia, Spain, Sweden, Switzerland, Tajikistan, Tunisia, Turkey, UK, Ukraine (Crimea, Kherson, Odessa), Uzbekistan, «Yugoslavia»; Oriental: China; Afrotropics: Madagascar, Tanzania, Yemen, St Helena (?introduced). =hamatus (Zetterstedt), 1843: 475 (Rhaphium); Becker, 1918: 281 (Syntormon); Grichanov, 2006: 199. Type locality: Scandinaviae (Lund, Scania, Ostrogothia, Thynaes, Norvegiae), Dania [Sweden, Norway, Denmark].
=pseudospicatus Strobl, 1899: 126; Becker, 1918: 281 (as a variation of S. pallipes); Negrobov, 1975: 659 (as a species); Grichanov, 2001: 182 (as a synonym of S. pallipes). Type locality: Spain: Algeciras. =uncitarsis Becker, 1902: 53; Becker, 1918: 281 (as a variation of S. pallipes); Negrobov, 1991: 55 (as a synonym of S. pseudospicatus). Type locality: Egypt: Fayȗm [=Al Fayyūm]; Mitte März. =immaculatus Santos Abreu, 1929: 414 (as a variation of S. pallipes); Negrobov, 1991: 55 (as a subspecies of S. pallipes). Type locality: Spain: Canary Is., La Palma. =longistylus Grichanov, 2001: 187 (as a subspecies of S. pallipes). Type locality: Madagascar: Manyakatompo. Material: 1♂, 1♀, Greece: Crete, Heraklion env., 7.VI.2000, Shamshev [VIZR]; 1♂, 1♀, [Ukraine: Crimea]: Karadag, Krym, 3-12.IX.1987, Tanasiichuk [ZIN]; 1♂, 1♀, Tajikistan: Sangvor, 10.VI.1977, Zlobin [ZIN]; 4♂, 1♀, Tajikistan: Kondara Gorge, Kvak tract, 12.VII.1977, Zlobin [ZIN]; 1♀, [Kyrgyzstan:] Osh Region, Bazar-Kurgan district, 12 km W Arslanbob, Ak-Terek, 1800 m, 27.VIII.1986, Tanasiichuk [ZIN]; 1♂, [Uzbekistan: Samarqand Region], Peyshambe / 12 [orange label; ex coll. A.P. Fedtshenko 1871; ZIN]. Remarks: The recent Catalogs (e.g., Negrobov, 1991; Yang et al., 2006) regarded S. pallipes and S. pseudospicatus to be different species. Grichanov (2001) considered them as colour phenotypes of the same species and did not find morphological differences between the two forms. They were often collected together; therefore, they cannot be subspecies. The subspecies S. p. longistylus Grichanov, 2001 known from Madagascar does not differ from both phenotypes in hypopygium morphology, but being very distinct in length ratio of antennomeres; otherwise it is closer to pseudospicatus phenotype. First record from Kyrgyzstan.
Syntormon pennatus Ringdahl, 1920: 25; Grichanov, 2006: 199. Type locality: Norway: Narvik. Distribution: Norway, Russia (Kabardino-Balkaria). Material (Fig. 2): 4♂♀, Kabardino-Balkaria, Ozen’, 43°13`N, 43°19`E, 13,15.VI.2001, Grichanov; 6♂, Kabardino-Balkaria, Bezengi, 43°10`N, 43°14`E, 18-19.VI.2001, Grichanov [ZIN]. Remarks: The material from Kabardino-Balkaria was directly compared with S. pennatus types deposited in the Lund Zoological Museum. See also remarks under S. turanicus.
Syntormon pumilus (Meigen), 1824: 53 (Porphyrops); Loew, 1857: 34-35 (Syntormon). Type locality: not given. Distribution: Afghanistan, Armenia; Austria, Belarus, Belgium, Bulgaria, Czech, Denmark, ?Egypt; Estonia, Finland, France, Germany, Great Britain, ?Greece, Hungary, Ireland, ?Israel, Italy, Latvia, Morocco, Norway, Poland, Romania; Russia (Astrakhan, Kabardino-Balkaria, Kaluga, Karelia, Krasnodar, Kursk, Leningrad, Lipetsk, Moscow, Murmansk, Novosibirsk, Pskov, Stavropol’, Vologda, Voronezh, Yakutia), Slovakia, ?Slovenia, Sweden, Spain (Canary Is.), Tunisia, Turkey, UK, Ukraine (Kherson, Odessa), «Yugoslavia»; «Middle Asia». (Some records may belong to S. denticulatus or S. triangulipes and should be confirmed). =longiseta (Zetterstedt), 1843: 471 (Rhaphium); Loew, 1850: 119 (Rhaphium); Becker, 1918: 282 (Syntormon); Grichanov, 2006: 199. Type locality: Sweden: «Ostrogothia, ad Haradshammar».
=pusillus (Zetterstedt), 1859: 5034 (Rhaphium); Becker, 1918: 282 (Syntormon); Grichanov, 2006: 199. Type locality: Sweden: Scania, Lindholmen. =pumilio (Zetterstedt), 1859: 5035 (Rhaphium); Becker, 1918: 282 (Syntormon). Type locality: Sweden: Scania, Lindholmen. =ibex Bezzi, 1903: 332 («Loew, olim in litt.»). =tridens (Becker), 1918: 253 (Xiphandrium); Negrobov, 1991: 56 (Syntormon). Type locality: «Warschau und Schlesien». =rufipes auctt., nec Meigen, 1824 (misidentification) (Collin, 1940: 268). Material: 1♂, Sweden: 7 km S Uppsala, 16.VI.1999, Grichanov [ZIN]; 1♂, [Russia]: Yashchera, Luzhskii Distr., Leningrad Region, 10.VII.1963, Stackelberg [det. A. Stackelberg as S. rufipes] [ZIN]; 1♂, [Russia]: Novosibirsk Region, Krasnozyorskoe, 21.VI.1988, Grichanov [ZIN]; 1♂, [Russia]: SW Yakutia, Biryuk River, 14 km upstream from Lena River, channel coast, on grasses at spruce forest, 16.VII.2008, A.K. Bagachanova [the Institute of Biological Problems of Cryolithozone, Siberian Branch of the Russian Academy of Sciences, Yakutsk]. Remarks: The species is firstly recorded from the Novosibirsk Region and Yakutia representing the easternmost point of its distribution.
Syntormon punctatus (Zetterstedt), 1843: 477 (Rhaphium); Becker, 1918: 276-277, 283 (as ?syn. of Syntormon denticulatus); Grichanov, 2006: 199. Type locality: Denmark. Distribution: Czech, Denmark, Germany, Norway, Poland, Russia (Leningrad, Moscow, Vologda), Sweden. =bisetosus Becker, 1918: 275; Negrobov, 1991: 56. Type locality: Poland: «Nimptsch in Schlesien». Material: 1♂, [Russia]: Sablino, Petrograd. Gub., 14.VIII.1924, Stackelberg [det. A. Stackelberg] [ZIN].
Syntormon rhodani Vaillant, 1983: 274. Type locality: France: «Alps, la vase de la cariçaie de Saint-Benoît sur les bords du Rhône». Distribution: France. Remarks: The species was incompletely described and illustrated, being a possible synonym to S. metathesis.
Syntormon samarkandi Negrobov, 1975: 659. Type locality: Uzbekistan: Kumak, Kattakurgan distr., Samarkand region. Distribution: ?Iraq, Uzbekistan. Remarks: Negrobov in his key to Palaearctic species (1975) misused the species name instead of S. giordanii Negrobov, 1974 (S. samarkandi was described with 5 strong black ventral bristles at base of fore femur, but was diagnosed in the key as bearing few long light hairs on the same place). Therefore, a record of S. samarkandi from Iraq (Olejnicek et al., 1995) may belong to S. giordanii.
Syntormon setosus Parent, 1938: 464 (described by Parent, 1927: 59 as Syntormon «espèce X»); Speight & Meuffels, 1989: 94. Type locality: France: «Dunes de Lion-sur- Mer (Calvados)». Distribution: Ireland, France, Italy, UK. Remarks: The species was originally described by a single female from northern France. Later females of S. setosus were reported from Italy and Ireland with its male remaining undescribed (see Speight & Meuffels, 1989). The latter authors supposed that S. setosus is a synonym of S. miki, distinguishing the two species by only variable colour characters.
Syntormon silvianus Pârvu, 1989: 57; Pârvu, 2000: 157; Grichanov, 2007: 70 (as synonym of S. monilis); Pârvu, 2009: 295 (restored). Type locality: Romania: Piatra Craiului Mountains, Southern Carpathians. Distribution: Romania. Remarks: See remarks under S. monilis.
Syntormon smirnovi Stackelberg, 1952: 403. Type locality: Tajikistan: Gissar Ridge, Varzob Gorge, Ziddy. Distribution: Kazakhstan, Kyrgyzstan, Tajikistan.
Syntormon spicatus (Loew), 1857: 33 (Rhaphium), nom. ressur.; Loew, 1857: 34-35 (Syntormon); Denninger, 1950: 45 (syn. of Syntormon fuscipes); here restored to species status. Type locality: Poland: «aus hiesiger Gegend» [= Meseritz]. Distribution: Poland. Remarks: The species was synonymised with S. fuscipes (von Roser) by Denninger (1950) who studied a male type of the latter species and compared it with the Figs 202204 provided by Becker (1918) for S. spicatus. However, Becker did not study Loew’s types, but compared the material in his hands with the original description of the species and found some differences between them, e.g. in presence of antennal scape setation in
S. spicatus sensu Loew and in hind tarsomere length ratio. Therefore, I consider Becker’s description belonging to a different species (S. fuscipes), and restore S. spicatus from synonymy, as I consider Denninger’s nomenclatural act to be based on Becker’s misidentification (see remarks under S. fuscipes and S. francoisi). The distribution of S. spicatus Loew must be confined to the type locality, because all subsequent records of the species were made for S. spicatus sensu Becker.
Syntormon subinermis (Loew), 1869: 290 (Synarthrus); Becker, 1918: 284 (Syntormon). Type locality: Slovakia: Losoncz [=Lučenec]. Distribution: Austria, Czech, France, Georgia, Germany, Hungary, Israel, Kyrgyzstan, Romania, Russia (Kabardino- Balkaria, Pskov, Vologda), Slovakia, Sweden, Tajikistan, Turkey, Uzbekistan. =asiaticus Negrobov, 1975: 663 (as a subspecies of S. subinermis). Type locality: Tajikistan: Gissar ridge. Material: 1♂, Russia: Pskov Region, Velikie Luki, 30.VI.1997, Grichanov [ZIN]; 1♂, Russia: Kabardino-Balkaria, 2 km E Psynodakh, 21.VI.2000, Grichanov; 1♂, Tajikistan, Sangvor, 10.VI.1977, Zlobin [ZIN]; 1♂, Uzbekistan: N Navoi, Karatau Mts., 20 km WSW Langar, Gumbaz, 27.V.1984, Tanasiichuk [ZIN]. Remarks: First record from Uzbekistan.
Syntormon submonilis Negrobov, 1975: 662. Type locality: Russia: North Caucasus, Fisht, Tybga. Distribution: Russia (Krasnodar). Here excluded from Kabardino- Balkaria. Remarks: The species was included in the key (Negrobov, 1975) as having dark spot at apex of hind femur in contrast to entirely yellow hind femur in S. monilis, though the last species was originally described with «hind femora dusky at the tip». See remarks under S. monilis.
Syntormon sulcipes (Meigen), 1824: 31 (Rhaphium); Becker, 1918: 284 (Syntormon). Type locality: not given. Distribution: Albania, Andorra, Austria, Belgium, Czech, France, Germany, Hungary, Romania, Russia (Karachai-Cherkessia), Switzerland, UK, «Middle Asia».
=oedicnemus (Loew), 1859: 15 (Synarthrus); Becker, 1918: 284 (Syntormon). Type locality: not given.
Syntormon tabarkae Becker, 1918: 285. Type locality: Tunisia: Tabarka. Distribution: France, Greece, Tunisia, «Yugoslavia».
Syntormon tarsatus (Fallén), 1823: 7 (Hydrochus); Zetterstedt, 1838: 713 (Dolichopus); Haliday, in Walker & al., 1851: 205 (Rhaphium); Kowarz, 1884: 109 (Syntormon); Becker, 1918: 286 (Syntormon); Grichanov, 2006: 183. Type locality: Sweden. Palaearctic: Austria, Belarus, Belgium, Czech, Denmark, Estonia, Finland, France, Germany, Hungary, Ireland, Latvia, Netherlands, Norway, Poland, Romania, Russia (Buryatia, Karelia, Kamchatka, Leningrad, Pskov), Slovakia, Sweden, UK, Ukraine (Kherson). =gratiosus (Meigen), 1824: 100 (Dolichopus); Loew, 1857: 35 (Syntormon); Becker, 1917: 140; 1918: 286 (Syntormon). Type locality: not given [Germany: «aus hiesiger Gegend»]. =obscurellus (Zetterstedt), 1838: 709 (Dolichopus) (misinterpretation of Fallén, 1823, p.p.); Becker, 1917: 150; 1918: 286 (Syntormon). Type locality: Sweden: «Lapponia Umensi, ad Tresunda». =palmipes (Meigen), 1824: 55 (Porphyrops); Becker, 1918: 286 (Syntormon). Type locality: not given. =vittatus (Macquart), 1834: 444 (Porphyrops); Becker, 1918: 286 (Syntormon). Type locality: France: Bordeaux. Material: 1♂, [Russia]: Yashchera, Luzhskii Distr., Leningrad Region, 16.VI.1963, Stackelberg [det. A. Stackelberg] [ZIN]; [Russia]: Leningrad Region, Vyborg distr., Bolshoi Berezovyi Is., 1.VIII.1979, Kandybina [ZIN]. Remarks: S. tarsatus along with the Nearctic S. palmaris (Loew, 1864) and Far Eastern
S. pseudopalmarae Negrobov et Shamshev, 1985 form rather distinct species group with their males having strongly widened and compressed laterally 2nd-4th segments of mid tarsus (see Negrobov & Shamshev, 1985).
Syntormon triangulipes Becker, 1902: 54. Type locality: Egypt: «Fayȗm [=Al Fay-yūm], in der Wüste bei Siala». Distribution: Azerbaijan, Egypt, France, Russia (Krasnodar), Spain, Tajikistan. Material: 1♂, [Russia]: Krasnodar env., Riv. Kuban’ (VNIIBZR), 4.VI.2000, Grichanov [ZIN]; 1♂, S Tajikistan, Dusti vil. [env.: 37.32°N, 68.82°E], summer 1984, Grichanov [ZIN]. Remarks: First records from Russia and Tajikistan.
Syntormon turanicus Stackelberg, 1927: 229. Type locality: «Turkestan, Chanatum, Kokand, montes Alaiensis, prope amniculum Kizilsu». Distribution: Kazakhstan, Kyrgyzstan, Uzbekistan. Material (Fig. 3): Holotype ♂, Alai / 22 [yellow label] / Typ. Syntormon turanicus Stackelberg sp.n. [ZIN]. Remarks: According to Stackelberg (1927), one male was collected by A.P. Fedtshenko at 22.VII.1871. The species differs from S. pennatus species concept in mid leg coloration only (see key below). Do they represent different species, subspecies or phenotypes of the same species, is the question for a future revision.
Syntormon zelleri (Loew), 1850: 121 (Rhaphium); Loew, 1857: 34-35 (Syntormon). Type locality: Italy: Sicilien. Palaearctic: Abkhazia, Austria, Azerbaijan, Belgium, Czech, France, Germany, Greece incl. Crete, Hungary, Iran, Ireland, Italy, Netherlands, Romania, Russia (Krasnodar), Slovakia, Spain, Switzerland, UK, “Middle Asia”. Material: 1♂, Abkhazia: 5 km SE Ritsa Lake, 10.VII.2004, Grichanov [ZIN].
Syntormon pilitibia sp. nov.
(Figs. 4–9)
Diagnosis. The new species belongs to a S. pumilus group of species (see below), being very distinct in having double ventral row of erect spinules along entire length of mid tibia. All other species of the group bear simple setulae on mid tibia, having no erect ventral spinules.
Type material. HOLOTYPE ♂, Israel: Menahamiya, 32.67 N, 35.57 E, 27.X.2911, N. Vikhrev [MZUM].
Etymology. From Latin pilus, tibia – ‘with haired tibiae’. Belongs to ventral row of fine erect spinules on mid tibia.
4
Figs. 4−5. Syntormon pilitibia sp. nov.:4 − habitus, 5 – antenna.
Description. Male: Head: Frons metallic blue-violet; face with black ground col-our, white pollinose; palpi and proboscis black; antenna black; scape setose dorsally; pedicel medianly with long projection, as long as scape; postpedicel tapering, 2.5 times longer than high; stylus dorsoapical, simple, with microscopic hairs; length ratio of scape to pedicel to postpedicel to stylomeres 1 and 2, 15/18/37/10/56; postoculars in single row, ventrally pale and dorsally black, and with some setae near cervix.
Thorax: Mostly greenish black, mesonotum with metallic violet-bronze reflections, metaepimeron with row of white hairs; katepimeron with 1-2 hairs; setae black; 6 dorsocentrals; 7-8 acrostichals short, uniseriate; median scutellars strong, laterals as weak side hairs, and with pair of fine pale hairs medianly along scutellar margin.
Legs: Coxae mostly yellow; mid coxa black except yellow apex; hind coxa brownish at base; fore coxa with short pale anterior hairs and some strong black apical setae; trochanters, femora and tibiae yellow; hind femur dirty yellow at apex; tarsi black from tip of basitarsus; hind basitarsus mainly brown; fore femur with 6-7 short black ventral setae at base, of which 2 basal setae longish, nearly as long as femur diameter; and with subapical posteroventral seta; fore tibia with 2 strong dorsal setae at 2/5 and at apex, 1 posterior apical seta, and with short but distinct anterodorsal setal serration along distal half; fore basitarsus simple; 2nd segment of the same tarsus slightly swollen, with small but distinct apicoventral projection; 2nd-4th segments each with 1-2 somewhat elongated dorsal setae; mid femur with 8-10 ventral setae in basal half, the longest setae nearly as long as diameter of femur; and with subapical pair of strong antero- and posteroventral setae; mid tibia with 2 antero- and 1 posterodorsal strong bristles in basal half, with strong apicoventral bristle, and with 2 ventral rows of erect hairs along entire length; the hairs as long as tibia diameter; mid tarsus simple; hind femur with strong anterior subapical seta, without posterior subapical seta; hind tibia with 3 antero- and 3 posterodorsal bristles, and with some short ventral setae; hind tarsus unmodified. Fore leg length ratio (from tibia to tarsomere 5): 71/32/12/9/9/9, mid leg: 92/39/19/14/9/11, hind leg: 125/29/30/21/13/14.
6
7
Figs. 6−7. Syntormon pilitibia sp. nov.:6 − fore tarsus, 7 – mid femur and tibia.
Wing: Hyaline; ratio of cross-vein dm-cu to apical part of CuA1, 24/38; lower calypter yellow, with fan of mainly black setae; halter yellow.
Abdomen: Mostly greenish black, and with black vestiture; 2nd tergum with large yellow lateral spots; 5th segment ventrally expanded to form hood for hypopygium; hypopygium black, with brown cerci.
Female: unknown.
Measurements (mm): Body length 2.5, antenna length 1.2, wing length 2.6, wing width 0.9.
8 9 Figs. 8−9. Syntormon pilitibia sp. nov.:8 − wing, 9 –hypopygium.
Syntormon macula mediterraneus ssp. nov.
(Figs. 10−13)
Diagnosis. The new subspecies differs from the current concept of S. macula in colour characters mainly. S. macula female was originally described with entirely dark abdomen, black hind coxa and dark spot on wing vein M (Parent, 1927). S. macula male was never illustrated, but was described (d’Assis Fonseca, 1949) and diagnosed in a key (d’Assis Fonseca, 1978) as follows: segments of front tarsus are uniformly decreasing in length towards apex, metatarsus is much shorter than combined length of remaining segments; middle and hind coxae are black, yellowish at apex; middle femur ventrally bears a fine bristle about middle, and a row of short black setulae from this bristle to base of femur; hind tarsus is simple, not noticeably thickened, uniformly dark; wing has a distinct brownish spot on apical section of discal vein. S. m. mediterraneus male corresponds to the description of female (Parent, 1927) and male (d’Assis Fonseca, 1949), but has 2nd, 3rd and 6th terga of abdomen largely yellow; hind coxa yellow; mid femur ventrally without a fine bristle and without a row of short black setulae; basal two segments of hind tarsus mainly yellow; wing with indistinct purplish spot on M flexion.
Type material. HOLOTYPE ♂, Israel: Baniass [Panyas], 13.VI.1982, A. Freidberg. PARATYPES: 4♂, Greece, Rhodes, Rhodes, 1-3.VII.1981, A. Freidberg [TAU].
Etymology. The subspecies is named after the Mediterranean Region.
Description. Male: Head: Frons metallic blue-violet; face with black ground col-our, white pollinose; palpi and proboscis black; antenna black; scape setose dorsally; pedicel medianly with long projection, longer than scape; postpedicel tapering, 2 times longer than high; stylus strictly subapical, simple, with microscopic hairs; length ratio of scape to pedicel to postpedicel to stylomeres 1 and 2, 12/16/35/3/43; postoculars in single row, pale, with upper 4-5 setae black, and with some setae near cervix.
10
11
12 13
Figs. 10−13. Syntormon macula mediterraneus ssp. nov.: 10 − habitus, 11 – antenna, 12 – wing,
13 – hypopygium.
Thorax: Mostly greenish black, mesonotum with metallic violet-bronze reflections, metaepimeron yellow, with row of white hairs; katepimeron with 2-3 white hairs; setae black; 6 dorsocentrals; 7-8 acrostichals uniseriate; median scutellars strong, laterals as weak side hairs.
Legs: Coxae mostly yellow; mid coxa brown-black except yellow apex; fore coxa with short pale anterior hairs and some strong black apical setae; trochanters, femora and tibiae yellow; tarsi brown from tip of 2nd segment; legs simple, without remarkable setae and cilia; fore femur with subapical posteroventral seta; fore tibia with 1 short dorsal seta at middle, and with short but distinct anterodorsal setal serration along distal half; mid femur with subapical pair of strong antero- and posteroventral setae; mid tibia with 3 antero- and 1 posterodorsal strong bristles, with 1 short ventral bristle, with 3 apicals; hind femur with strong anterior subapical seta, 1 posteroventral subapical seta; hind tibia with 3 antero- and 5 posterodorsal bristles, and with some short ventral setae. Fore leg length ratio (from tibia to tarsomere 5): 92/50/24/18/10/10, mid leg: 130/59/30/20/12/10, hind leg: 167/34/35/22/17/13.
Wing: Hyaline, wing small dark spot on apical section of M1+2, sometimes poorly discernible; ratio of cross-vein dm-cu to apical part of CuA1, 30/43; lower calypter yellow, with fan of black setae; halter yellow.
Abdomen: Mostly yellow, and with black vestiture; 1st, 3rd and 6th terga partly, 4
5th
terga mostly brown-black with greenish tinge; 5th segment ventrally expanded to form hood for hypopygium; hypopygium yellow, with yellow cerci. Female: unknown.
Measurements (mm): Body length 3.0, antenna length 1.0, wing length 3.1, wing width 1.0.
Doubtful species Syntormon decoratus (Haliday), 1832: 353 (Plectropus); Becker, 1918: 286 (Syntormon, unrecognized). Type locality: Ireland: Holywood.
Species excluded from Syntormon Sympycnus praeteritus Parent, 1929: 9; Yang et al., 2006: 507 (Syntormon). Type locality: Germany: Rosenhof. Remarks: The species was described by a single female from Germany. According to the original description, Sympycnus praeteritus along with Syntormon bicolorellus and Sympycnus simplicipes Becker, 1908 represents one more intermediate form sharing features of both Sympycnus and Syntormon (=Bathycranium) that was noted by Parent (1929) himself. Yang et al. (2006) placed the species in the latter genus, giving no any explanation. Despite the strongly bulging face, it would be better to keep S. praeteritus in Sympycnus until the type will be studied. The broad bulging face is a female secondary sexual character found in Telmaturgus Mik, 1874, Parasyntormon Wheeler, 1899, monotypic genera Hercostomoides Meuffels et Grootaert, 1997, and Ceratopos Vaillant, 1952, in some species of Sympycnus Loew, 1857 (e.g. S. simplicipes) and Teuchophorus Loew, 1857 (e.g. T. longifrons Bickel, 1983, and T. queenslandicus Bickel, 1983) in addition to Syntormon. Rhaphium rufipes Meigen, 1824: 30; Becker, 1918: 277-278 (unrecognized); Parent, 1925: 42 (syn. of Syntormon pumilus); Collin, 1940: 268 (unrecognized species of ?Rhaphium). Type locality: not given.
Key to West-Palaearctic species and species groups of Syntormon (males) Remarks: S. luteicornis and S. setosus known from females only, as well as subspecies of known species (S. macula mediterraneus and S. subinermis asiaticus) and phenotypes and subspecies of S. pallipes are not included in the key. As follows from my remarks (see above), I consider colour characters used in the previous keys to be very variable in the genus and inappropriate for distinguishing Syntormon species. It seems also that the length ratio of antennomeres and ratio of length and width of postpedicel are variable to a certain extent, as well as the length and number of ventral setae on femora. In addition, some species were incompletely described or poorly illustrated. Therefore, I have compiled a new key based mainly on male secondary sexual characters, indicating the following species groups that must be revised in the future:
Syntormon fuscipes group of species: simple fore tarsus and strong basoventral projection on hind basitarsus, bearing modified setae on apex of that projection: francoisi, fuscipes, spicatus; S. valae Negrobov et Zhilina, 1986 described from Mongolia also belongs to this species group (see Grichanov, 2001);
Syntormon monilis group of species: modified tarsomeres of fore and mid tarsus and strong basoventral projection on hind basitarsus, bearing modified setae on apex of that projection: monilis, silvianus, submonilis;
Syntormon sulcipes group of species: simple tarsi except for the presence of divergent ventral setae on hind basitarsus, of which basal seta is stronger and directed basally: bulgariensis, obscurior, sulcipes; S. siplivinskii Negrobov, 1975 (Buryatia) is related to this species group;
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Syntormon pennatus group of species: the ventral setae on hind basitarsus as in species of sulcipes group, but plumose dorsally and ventrally hind tibia and plumose dorsally hind tarsus: pennatus, turanicus;
Syntormon pumilus group of species: shortened fore tarsomeres, with at least 2nd segment of same tarsus swollen; more or less elongate basoventral setae or cilia on fore and mid femora:
– metathesis subgroup of species: no apical lobe on 2nd segment of fore tarsus: metathesis, rhodani.
– pumilus subgroup of species: distinct apical lobe on 2nd segment of fore tarsus; abdomen entirely dark (pumilus, iranicus) or abdomen with 2nd-3rd segments partly yellow (giordanii, samarkandi, triangulipes).
1. Hind basitarsus simple..................................................................................................2
– Hind basitarsus bearing processes, spines, leaf-like or long setae .................................9
2. Fore tarsus having segments regularly decreasing in length towards apex; basitarsus much shorter than combined length of remaining segments..........................................3
– Fore tarsus with shortened 2nd-4th segments, with at least 2nd segment of same tarsus swollen; fore and mid femora with more or less elongate basoventral setae or cilia (pumilus group of species) .............................................................................................7
3. Antenna mostly or partly yellow; abdominal tergites extensively yellow; legs entirely yellow and simple; 2.5-3.0............................................................................bicolorellus
– Antenna entirely dark.....................................................................................................4
4. Wing with small dark spot on M1+2 just before middle of its distal part; legs simple; mid femur with at most one fine bristle about middle; mid tibia without long ventral setae; 3.0-4.0 ........................................................................................................macula
– Wing clear; either mid femur or hind tibia bearing long ventral setae...........................5
5. Mid femur without long ventral setae; hind tibia with posteroventral row of 8-10 long erect setae (Fig. 11); 5.0...................................................................................... cilitibia
– Mid femur with long ventral setae; hind tibia without long erect setae .........................6
6. Mid femur with 2-3 long ventral setae; hind tarsus distinctly thickened; 4.0-4.5....miki
– Mid femur with 12-14 long ventral setae; hind tarsus not thickened; 3.0............codinai
7. 2nd segment of fore tarsus slightly enlarged, without apical lobe; 3.2-4.5 ...................... ....................................................................................... metathesis subgroup of species
– 2nd segment of fore tarsus with distinct apical triangular prolongation..........................8
8. Mid tibia with double ventral row of erect hairs along entire length; the hairs as long as tibia diameter ..................................................................................................pilitibia
– Mid tibia with simple setulae, without erect ventral spinules; 1.7-3.0............................. ........................................................................................... pumilus subgroup of species
9. Hind tibia strongly thickened, bearing rows of long setae dorsally and ventrally ......10
– Hind tibia simple or slightly thickened, usually without rows of long setae ...............11
10. Hind tibia with simple setae, longish ventrally; hind tarsus without dorsal plumation; hind basitarsus with basoventral tubercle bearing leaf-like branched appendix;
2.5 ......................................................................................................................smirnovi
– Hind tibia plumose dorsally and ventrally; hind tarsus plumose dorsally; hind basitarsus bearing same ventral setae as sulcipes group of species; mid femur and tibia mainly yellow (pennatus: Fig. 12) or blackish except yellow knees (turanicus: Fig. 13); 2.5 ............................................................................................... pennatus group of species
11. Some apical segments of mid tarsus widened and flattened .....................................12
22
– Mid tarsus simple.........................................................................................................14
12. 2nd-4th segments of mid tarsus strongly widened and compressed laterally, black; hind tarsus black; hind tibia at apex thickened, black; hind tarsus black; hind basitarsus with long curved ventral seta; postpedicel 3 times longer than high at base; 3.0. ............................................................................................................................tarsatus
– 4th and 5th segments of mid tarsus widened..................................................................13
13. 4th and 5th segments of hind tarsus widened; hind tibia thickened and slightly curved; hind basitarsus without spiniform ventral process, with 3 setae of unequal length; fore tarsus simple; postpedicel 1.2 times longer than high at base; 3.6-3.7.............. latitarsis
– 4th and 5th segments of hind tarsus not widened; hind tibia not thickened; hind basitarsus with long pointed basoventral process; fore tarsus ciliated, with shortened 2nd-4th segments; 2.5-3.0 ......................................................................monilis group of species
14. Hind basitarsus with long simple ventral setae .........................................................15
– Hind basitarsus with short hook-like curved setae, with leaf-like appendages or with process .........................................................................................................................17
15. Fore femur bearing long ventral seta at base; 1st and 2nd segments of hind tarsus each with 1 erect ventral seta at about middle, that on 2nd segment longer and square-ended; 2.0 .........................................................................................................................filiger
– Fore femur without long ventral seta; 2nd segment of hind tarsus without ventral seta.16
16. Hind basitarsus with 2 strong ventral setae of equal length; 3.0................... punctatus
– Hind basitarsus with 2 divergent ventral setae of unequal length; hind tibia laterally compressed and markedly club-shaped in lateral view; 3.0-3.3sulcipes group of species
17. Hind basitarsus with leaf-like or worm-like ventral appendages, without unguiculate ventral spines or process ..............................................................................................18
– Hind basitarsus with unguiculate ventral spines or with process.................................19
18. Hind basitarsus with 1 leaf-like pedunculate ventral appendix; 2.5-3.0 ..... subinermis
– Hind basitarsus swollen at base ventrally, here with pair of yellowish oval leaf-like appendages surrounded by fine black setulose hairs; 3.0-3.5 ................................zelleri
– Hind basitarsus with about 5 worm-like ventral appendages on basal swelling; 3.0 ....... .........................................................................................................................mutillatus
19. Hind basitarsus with basoventral or midventral pointed process or tubercle bearing bunch of modified setae ...............................................................................................20
– Hind basitarsus with only two bare ventral hooks in basal half, without modified setae or process .....................................................................................................................22
20. Hind basitarsus swollen ventrally at middle; the swelling having a small tubercle bearing leaf-like pedunculate bilobate appendix in addition to rigid process terminating with two short hooks; 4.0............................................................................ tabarkae
– Hind basitarsus swollen ventrally in basal 1/5, excavated ventrally in middle 1/3; the swelling having pointed process ..................................................................................21
21. Hind basitarsus with pointed process bearing 3 subapical worm-like appendages and 2 short thick apical setae; 2.5 ........................................................................abbreviatus
– Hind basitarsus with pointed process bearing apical bunch of 2-3 worm-like and some shorter thick apical setae; 2.1-3.25 ..........................................fuscipes group of species
22. Postpedicel 3-3.5 times longer than high, nearly 2 times longer than stylus; mid femur without strong ventral setae; hind basitarsus with strong, fused at base ventral hooks; 3.0-3.75 ...................................................................................................pallipes
– Postpedicel at most 1.5-2 times longer than high, shorter than stylus; mid femur with row of ventral setae; hind basitarsus with weak, spiniform, distinctly separated ventral setae .............................................................................................................................23
23. Abdomen brownish yellow, with dark posterior margins of tergites; hind basitarsus with 2 slightly curved ventral setae; 4.0-6.0 ........................................................ aulicus
– Abdomen entirely dark; hind basitarsus with 2 strong curved ventral spines..............24
24. Hind basitarsus with ventral spines at middle; lower calyper with black cilia; 5.0-5.5 .......................................................................................................................freymuthae
– Hind basitarsus with ventral spines at base..................................................................25
25. Postpedicel 1.5-2 times longer than high; lower calyper with white cilia; 2.5-4.0 ....... ..................................................................................................................... denticulatus
– Postpedicel as long as high; lower calyper with ?white cilia; 2.0-3.0........... brevicornis
Acknowledgements
The author expresses sincere gratefulness to Dr. A. Ozerov (MZUM), Dr. E. Nartshuk (ZIN) and Dr. A. Freidberg (TAU) for their kindness in furnishing an opportunity to study the collections of their museums. Dr. S.Yu. Storozhenko (Vladivostok) has made some useful comments. This paper was partly supported by the grant of the Russian Foundation for Basic Research N 11-04-01051-a to Oleg P. Negrobov.
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AFROTROPICAL SPECIES OF THE GENUS ASYNDETUS LOEW (DIPTERA: DOLICHOPODIDAE) WITH NOTES ON SOME PALAEARCTIC AND ORIENTAL SPECIES
Igor Ya. Grichanov
All-Russian Institute of Plant Protection (VIZR), Podbelskogo 3, St.Petersburg, Pushkin, 196608, Russia. E-mail: grichanov@mail.ru
The genus Asyndetus Loew, 1869 in the Afrotropical Region is reviewed. It comprises 14 species including five new species and two doubtful species described by females. The genus is differentiated from other Diaphorinae by the following combination of characters: vein M broken and displaced; upper part of proepisternum with 1–4 fine setae; acrostichals present, biserial; male segment 8 with strong projecting setae. New species A. congensis sp. n. from D.R. Congo, A. savannensis sp. n. from D.R. Congo and Senegal, A. madagascarensis sp. n. from Madagascar,
A. namibiensis sp. n. from Namibia and A. pseudoseparatus sp. n. from Gambia are described. A key to the Afrotropical species of Asyndetus is provided, as well as new records for some Afro-tropical, Palaearctic and Oriental species. Lectotype and paralectotypes are designated for A. albipalpus Loew, 1871.
KEY WORDS: Dolichopodidae, Asyndetus, Afrotropical, Palaearctic, India, new species, new records, key.
И.Я. Гричанов. Виды рода Asyndetus Loew (Diptera: Dolichopodidae) тропической Африки с заметками о некоторых палеарктических иориентальных видах
Дан обзор афротропических видов рода Asyndetus Loew, 1869, который содержит 14 видов, включая пять новых для науки и два сомнительных вида, описанных по самкам. Род отличается от других родов подсемейства Diaphorinae следующими признаками: жилка крыла M в вершинной части с разрывом; проэпистернум с 1–4 щетинками в верхней части; акростихальные щетинки двурядные; 8-й сегмент брюшка с крепкими длинными щетинками. Описаны и иллюстрированы A. congensis sp. n. из Д.Р. Конго, A. savannensis sp. n. из Д.Р. Конго и Сенегала, A. madagascarensis sp. n. с Мадагаскара, A. namibiensis sp. n. из Намибии, A. pseudoseparatus sp. n. из Гамбии. Приведены определитель афротропических видов Asyndetus и новые указания для известных видов из тропической Африки, Индии и Палеарктики. Обозначены лектотип и паралектотипы для A. albipalpus Loew, 1871.
Всероссийский научно-исследовательский институт защиты растений (ВИЗР), шоссе Подбельского, 3, Санкт-Петербург, Пушкин, 196608, Россия.
Introduction
The genus Asyndetus Loew, 1869 belongs to the subfamily Diaphorinae. There are more than 100 species of the genus (Grichanov, 2003–2013) reported from all zoogeographical regions of the Earth. In the Old World, it is quite diverse in the tropical and subtropical belts. Negrobov (1973) published the last review of Palaearctic Asyndetus species. Subsequently Bickel (1996) redescribed the genus, and Grootaert & Meuffels (2002) defined the A. latifrons species group. Oriental and Australasian faunas of the genus were treated by Meuffels & Grootaert (1993), Grootaert & Meuffels (2002), Bickel (1996), Zhang & Yang (2003), Wang et al. (2007).
Curran (1926a, b) was the first who described Asyndetus species from the Afro-tropical Region (three species from South Africa). Few data, including descriptions of new species, have subsequently been published on the genus. Parent (1929a, 1930, 1937) and Séguy (1950) described 5 new species from DR Congo, Madagascar, Niger and the Hala'ib Triangle region. Grichanov (2000, 2012) found the South African A. virgatus Curran in Namibia and Sierra Leone; the Madagascan A. decaryi Parent was
28
recorded from South Africa (Grichanov & Urban, 2009) and Kenya (Grichanov et al., 2011a). A. latifrons (Loew) widely distributed in the Palaearctic and Oriental Regions was firstly reported from the Afrotropics (Gabon) by Grichanov (2011). There were no suitable keys to the Afrotropical species.
Material and methods
The holotypes and paratypes of the new species and other material cited are housed at the All-Russian Institute of Plant Protection, St. Petersburg, Russia (VIZR), the Finnish Museum of Natural History, Helsinki, Finland (MZH), the Hungarian Natural History Museum, Budapest, Hungary (HNHM), the Museum of Zoology, Lund University, Lund, Sweden (MZLU), the Natal Museum, Pietermaritzburg, South Africa (NMSA), the National Museum, Bloemfontein, South Africa (BMSA), the National Museum of Natural History, Paris, France (MNHN), the Natural History Museum, London, United Kingdom (BMNH), the Natural History Museum of Denmark (ZMUC), the Naturhistoriska Riksmuseet, Stockholm, Sweden (NHRS), the Royal Belgian Institute of Natural Sciences, Brussels, Belgium (IRSNB), the Royal Museum for Central Africa, Tervuren, Belgium (RMCA), the Zoological Institute of the Russian Academy of Sciences, St. Petersburg (ZIN), the Zoological Museum of Moscow State University, Russia (ZMU), the Zoological Museum of Tel-Aviv University, Israel (TAU), and the Zoologisches Museum, Universität Kiel, Kiel, Germany (ZMUK).
Specimens were studied and illustrated with a ZEISS Discovery V–12 stereomicroscope and an AxioCam MRc5 camera. Morphological terminology and abbreviations follow Cumming & Wood (2009). The relative lengths of the podomeres should be regarded as representative ratios and not measurements. Body length is measured from the base of the antenna to the tip of abdominal segment 8. Wing length is measured from the base to the wing apex. Male genitalia were macerated in 10% KOH. Figures showing the male genitalia in lateral view are oriented as they appear on the intact specimen, with the morphologically ventral surface of the genitalia facing up, dorsal surface down, anterior end facing right and posterior end facing left. Information on world distribution for known species follows Grichanov (2003–2013).
Systematics
Family Dolichopodidae Subfamily Diaphorinae Genus Asyndetus Loew
Asyndetus Loew, 1869: 34. Type species: Asyndetus ammophilus Loew, 1869, by designation of Coquillett, 1910: 511.
Meringopherusa Becker, 1902: 56; Strobl, in Czerny, Strobl, 1909: 189. Type species: Meringopherusa separata Becker, 1902, by designation of Dyte, 1975: 247.
Diagnosis. This cosmopolitan genus is defined (together with Cryptophleps Lichtwardt, 1898) by the synapomorphy of the broken and displaced vein M which readily distinguishes it from the related and probably ancestral genus Diaphorus Meigen, 1824. Small to medium-sized flies (body length 1.5-4.5 mm); upper part of proepisternum with 1–4 fine setae; acrostichals present, biserial; male segment 8 with four strong projecting setae (Grichanov, 2011). A thorough re-description of all the characters can be found in Bickel (1996). See also the definition of A. latifrons species group by Grootaert & Meuffels (2002).
Remarks. The closely related genus Cryptophleps was originally distinguished from Asyndetus by lacking wing cross-vein dm-cu (Lichtwardt, 1898), and was later distinguished from Asyndetus by lacking mesonotal acrostichal setae (Lamb, 1922; Parent, 1938), hypopygial macrochetae (Negrobov, 1973; Grootaert & Meuffels, 1987) or upper proepisternal setae (Bickel 2005), by reduced number of notopleural and supra-alar setae (Negrobov, 1973). All these characters are quite variable within the Asyndetus, Cryptophleps and other genera of Diaphorinae, correlating mainly to the size of species. By now the presence or absence of acrostichal setae is the only appropriate character for distinguishing the two genera (Capellari & Grichanov, 2012).
Composition. The genus numbers more than 100 mostly Holarctic species, widely distributed across arid zones of Palaearctic and Afrotropical Regions. Thirteen species are recorded from Afrotropical Region.
Review of Afrotropical species
Asyndetus albifacies Parent, 1929a: 46. Type material examined: Paratypes: 2♂, M. Halaib, 21.I.[19]29 / Coll. Efflatoun, Egypte / Asyndetus albifacies Par. ♂, cotype / paratype [MNHN]. Material examined: 1♂, Saudi Arabia: Bahara, March 77, Dr. Buttiker / 7 [BMNH]; 1♂, 2♀, Saudi Arabien, W. Buttiker / Bahara, 24.3.76 [BMNH]; 1♂, Saudi Arabien, W. Buttiker / Selouly's Farm, 30.10.75 [BMNH]; 2♂, Israel: Eilat env., ~29.57°N, 34.97°E, 25.X.2011, N. Vikhrev [ZMU]. Distribution: Type locality: Mt. Halaib. Afrotropical: Hala'ib Triangle (Egypt–Sudan border); Palaearctic: Israel, Saudi Arabia. New for Israel and Saudi Arabia.
Asyndetus albifrons Parent, 1929a: 45. Type material examined: Paratypes: 2♂, 2♀, Bir Abrag, South Eastern Desert, 3.III.1928 / Coll. Efflatoun, Egypte / Asyndetus albifrons Par. ♂, cotype / paratype [MNHN]; 1♀, same labels [RMCA]. Distribution: Type locality: Bir Abrag (South Eastern Desert). Palaearctic or Afrotropical: Egypt (close to Hala'ib Triangle); Palaearctic: Iraq.
Asyndetus amaphinius Séguy, 1950: 275. Type material examined: Holotype ♂: Monts Tarraouaji, 900 m, 8-12-IX / IFAN-1947,
L. Chopard, A. Villiers / Type [red label] / Asyndetus amaphinius Type, E. Séguy vid. 48 [MNHN]. Distribution: Type locality: Niger: Aïr, Monts Tarraouaji. Afrotropical: Niger.
Asyndetus crassitarsis Curran, 1926b: 410. Distribution: Type locality: South Africa: Transvaal, Kaapmuiden. Afrotropical: South Africa.
Asyndetus decaryi Parent, 1929b: 187 (in key); 1930: 100 (description). Type material examined: Holotype ♂: Madagascar: Prov. d’Ananalava, Maromandia, B. Decary, 1922 / Type [red label] / Asyndetus decaryi n. sp. ♂, O. Parent det. [MNHN]. Material examined: 3♂, 3♀, South Africa: Cape Province, 10 km S of Koomlanskloof, 32°40'S, 19°02'E, Malaise-trap, marshy meadow at riverside, 5-7.X.1994, M. Söderlund
[ZMLU]; 1♂, RSA: Cape Prov. 9 km NW Worcester, 33°37'S, 19°22'E, 9.X.1994, Loc. 11, leg. R. Danielsson [ZMLU]; 1♂, South Africa: Cape Province, Umngazi Mouth, 3129Da, 20.X.1972, M.E. Irwin, 3 to 10 m, coastal dunes [NMSA]; 1♂ (in alcohol), Madagascar: Foulpointe, 2.XI.1991, A. Pauly col., plage, bac jaune [IRSNB]; 2♂ (in alcohol), Madagascar: Foulpointe, X.1993, A. Pauly col., forêt, lagune, Piège Malaise [IRSNB]; 6♂, 2♀ (in alcohol), Madagascar: Tam., Foulpointe, 2.XI.1991, A. Pauly col., plage [IRSNB]; 3♂ (in alcohol), Madagascar: 25 km W. Morarano-chrome, XI.1991, forêt, bac jaune, A. Pauly [IRSNB]; 2♂ (in alcohol), Madagascar: Manakambahiny, Atn., I.1991, A. Pauly [IRSNB]. Distribution: Type locality: Madagascar: province d’Analalava, Maromandia. Afro-tropical: Kenya, Madagascar, South Africa.
Asyndetus latifrons (Loew), 1857: 46) (Diaphorus); Loew, 1869: 298 (Asyndetus); Negrobov: 1973: 165 (Fig. 25); Pârvu, 1995: 298 (Fig. 5); Grootaert & Meuffels, 2002: 43 (Fig. 7); Grichanov et al., 2011b: 26 (Fig. 8), 33 (Fig. 78). Material examined: Afrotropical: 1♂, 1♀, Kenya: Mombasa, 13.VIII.1983, A. Freidberg [TAU]; 1♂, [DR Congo:] Congo-belge, Bambesa, 22.VIII.1938, J. Vrydagh [IRSNB]; 1♂, (in alcohol), Gabon: Kougouleu, IX.1984, A. Pauly, Piège Malaise [IRSNB]; 4♂, 4♀ (in alcohol), Gabon: Voleu-Ntem, Assok-Ngum, 21.II.1986, A. Pauly, bac jaune, coupe forestière [IRSNB]; 38♂♀ (in alcohol), Gabon: Libreville, IX.1984, A. Pauly [IRSNB]; 16♂♀ (in alcohol), Gabon: Owendo, 9.XII.1985, A. Pauly, dunes littorales, bacs jaune [IRSNB]; Palaearctic: 1♂, [Russia:] Krasnodar Territory, Arkhipo-Osipovka, 14-26.VI.1992 / peach orchard, Grichanov; 1♂, Krasnodar Terr., Gelendzhik, env. vill. Betta, stream r. Betta, on stones, 11.VII.2006, Volfov, Talashinskii [VIZR]; 1♀, Russia: N Osetia-Alania, Alagir env., 5.VII.2000, I. Grichanov [VIZR]; 1♀, Russia: Adygea, Maikop, Vostochnaya str., garden, 4.IX.2009,
K. Tomkovich [ZMU]; 2♂, 1♂, [Russia:] Belgorod Region, Borisovka vil., 5.VII.2001,
D.D. Kostrov [ZMU]; 1 ♂ [size 3 mm], Yermasoyia Riv., Cyprus, coll. G. Mavromoustakis, 13.7.56 / R.I.Sc.N.B. 24236, Coll. M. Bequart [IRSNB]; 1♀, Turkey: Adapazari reg., Karasu env., 27.VIII.2009, N. Vikhrev, N. Dvoretskaya [ZMU]; 1♂, Turkey: Zonguldak reg., Alapli env., 41.14°N, 31.36°E, 19-20.VI.2010, N. Vikhrev [ZMU]. Distribution: Type locality: Poland: “Schlesien”. Afrotropical: DR Congo, Gabon, Kenya; Palaearctic: Austria, Belgium, Bulgaria, Cyprus, Czech, Estonia, France, Germany, Hungary, Italy, N Kazakhstan, Netherlands, Poland, Romania, Russia (Adygea, Krasnodar, Leningrad, N Osetia-Alania, Voronezh, S Ural), Slovakia, Spain, Syria, Turkey; Oriental: Bangladesh, China, India, Pakistan, Philippines, Thailand. New for Kenya, DR Congo, Cyprus, some Russian regions (Adygea, Belgorod, N Osetia-Alania).
Asyndetus virgatus Curran, 1926a: 34; Grichanov, 2012: 2 (Figs. 1–2) Type material examined: Holotype ♂, Paratype ♂: Middelburg [25º47'S:29º28'E], 13.2.[19]25, H.K. Munro [NMSA]; Paratype 1♀: East London, 9.3.25, H.K. Munro [NMSA]. Material examined. 2♂, Natal: Weenen, IV.1924, H.P. Thomasset / Pres. by Imp. Inst. Ent. Brit. Mus. 1932-338 / Asyndetus virgatus Curr., det. C.E. Dyte; 1 ♀, Natal: Weenen, 2840 ft., I.1924, H.P. Thomasset, Thorn country [other labels are the same; BMNH]; 2♂, 3♀, South Africa: Natal, Ashburton, 15 km SE of Pietermaritzburg,
II.1977, J.G.H. Londt, Malaise in Grassland [NMSA]; 1♂, South Africa: Natal, Lynn-field Park, 13 km SE of Pietermaritzburg, 29°41’S, 30°29’E, Acacia thornveld area, 31.III.1989, A.E. Wittington [NMSA]; 1♂, South Africa: Natal, 15 km SE Ingwavuma, 2732Aa, 21.II.1979, J.G.H. Londt, Bushy area with big trees ex Malaise trap [NMSA]; 1♂, South Africa: KwaZulu-Natal, Royal Natal National Park, Mahai River, 28°41.386’S, 28°56.288’E, 17–18.II.2010, Malaise trap (1), straddling Mahai River,
A.H. Kirk-Spriggs [BMSA]; 25♂♀, South Africa: Free State, Brandfort, Florisbad Res. Stat., 28°46.039’S, 26°04.234’E, 4–6.IV.2009, Malaise traps, Acacia savanna, A.H. Kirk-Spriggs [BMSA]; 1♂, South Africa: KwaZulu-Natal, Ndumo Game Reserve, Shokwe area, 26°52.125’S, 32°13.731’E, 30.XI–4.XII.2009, Malaise traps, Ficus forest, A.H. Kirk-Spriggs [BMSA]; 1♀, South Africa: KwaZulu-Natal, Ndumo Game Reserve, main road, 26°54.288’S, 32°17.974’E, 4–8.XII.2009, Malaise traps, sand [forest] & broad-leafed deciduous forest, A.H. Kirk-Spriggs [BMSA]; 1♀, South Africa: E. Cape, Tsitsikamma National Park, Plaatbos Nature Reserve, 33°59.283’S, 23°55.275’E, 20–22.I.2009, Malaise trap, Oubrug, Storms River margin, indigenous forest, A.H. Kirk-Spriggs & S. Otto [BMSA]; 8♂, 8♀ [in alcohol], [Ivory Coast:] C.I., Komandimi, NP Comoe, Camp Univ. Würzb., 8°45’ N, 3°49’ W, G-Schale, 13-31.I.1998, leg. S.Hilger [ZMUK]; 2♂, 1♀ (in alcohol), Gabon: Ntoum, X.1984, A. Pauly, Piège Malaise, Pâturage [IRSNB]; 2♀ (in alcohol), Gabon: Booué, IX.1984, A. Pauly, Piège Malaise [IRSNB]; 5♂, (in alcohol), Gabon: Ntoum, X.1985, A. Pauly, , bac jaune, plantation sur brûlis [IRSNB], 36♂♀ (in alcohol), Gabon: Lastourville, VIII-X.1984, A. Pauly, Piège Malaise [IRSNB]; 4♂, 5♀ (in alcohol), Guinea-Bissau: Bafatá, Bafatá, 810.IX.2004, N. Jönsson [ZIN]; 1♂, [DR Congo:] Congo-belge, Rutshuru, 30.XI.1937, J. Ghesquière [IRSNB]; 1♂, [DR Congo:] Congo-belge, Kivu, Rutshuru (riv. Fuku), 1250 m, 6.VII.1935, G.F. de Witte: 1682 [IRSNB]; 1♂, [DR Congo:] Congo Belge, P.N.G., Miss. H. De Saeger, II/gd/4, 24.VII.1952, H. De Saeger, 3864 [RMCA]; 1♂, [DR Congo:] Congo Belge, P.N.G., Miss. H. De Saeger, II/gd/4, 18.IX.1952, H. De Saeger, 4077 [RMCA]; 1♂, [DR Congo:] Congo Belge, P.N.G., Miss. H. De Saeger, II/gd/8, 10.IV.1952, H. De Saeger, 3313 [RMCA]; 1♂, [DR Congo:] Congo Belge, P.N.G., Miss. H. De Saeger, II/fd/17, 3.I.1952, H. De Saeger, 2991 [RMCA]; 1♂, Kenya: Tiwi, Beaches, 4 14 S, 39 36 E, 14-23.VIII.1975, B. Petersen [ZMUC]; 1 ♂, Ghana: Kumasi, No. 13, 18-20.VI.1965, leg. Endrody-Younga [HNHM]; 1♂, Botswana: SE district, Gaborone city, National Museum garden, light trap, 8-10.XI.1993, Bert A.V. Viklund [NHRS]; 1♂, Malawi: Kasungu Nat. Park, Lifupa Camp, 1333Aa, 9-10.XII.1980, 1000 m, Stuckenberg & Londt, Brachystegia [NMSA]; 1♂, 1♀, Yemen: Ma’rib, Ma’rib, 24.IV.1992, R. Linnavuori [MZH]. Remarks: A male from Yemen differs in the presence of 6 strong bristles (rather than 4) on segment 8 and elongate setulae on anteroventral and anterodorsal surfaces of hind tibia, about as long as diameter of tibia. Surstyli of studied specimens are variable to a certain extent in length and setulation. Distribution: Type locality: South Africa: Mpumalanga: Middelburg. Afrotropical: Botswana, DR Congo, Kenya, Gabon, Ghana, Guinea-Bissau, Ivory Coast, Malawi, Namibia, Sierra Leone, South Africa, Yemen. New for Botswana, DR Congo, Kenya, Gabon, Ghana, Guinea-Bissau, Ivory Coast, Malawi, and Yemen.Asyndetus pseudoseparatus sp. nov.
(Fig. 1)
Diagnosis. A. pseudoseparatus differs from other Afrotropical species of the genus in lacking cross-vein dm-cu, distinctly broken M1+2, simple tarsi bearing claws and small pulvilli. It is very close to A. separatus in external morphology, differing from the latter in the presence of claws and small pulvilli on at least anterior four tarsi, in absence of strong ventral setae on mid and hind femora. A. separatus male has enlarged tarsal pulvilli, about half as long as segment 5, lacking claws on fore and mid tarsi and having one claw on hind tarsus. Male genitalia of the two species are very close to each other except for the shorter mid-dorsal seta on ventral lobe of surstylus in A. separatus, half as long as figured for A. pseudoseparatus (Fig. 1). In addition, an examined A. separatus male from Egypt has asymmetrical epandrial lobes (see below).
Type material. HOLOTYPE ♂, Gambia: Kotu stream about 3 km SW Bakau, in vegetation and fresh water, 23.II.1977, UTM 28PCK16-88, Loc. No. 3 / Lund Univ., Syst. Dept., Sweden Gambia/Senegal. Febr.-March 1977, Cederholm - Danielsson -Larsson - Mirestrom - Norling -Samuelsson [ZMLU]. PARATYPES:1 ♀, same labels; 1 ♀, Gambia: about 1 km E Yoga Jula milestone 105, 2.III.1977, Loc. No. 13, UTM 28PDK28-83 / Lund Univ., Syst. Dept., Sweden Gambia/Senegal. Febr.-March 1977, Cederholm - Danielsson - Larsson - Mirestrom - Norling - Samuelsson [ZMLU].
Etymology. The species is named for its similarity with Asyndetus separatus Becker.
Description. Male: Head: frons bronze-black, grey pollinose; face black, densely whitish pollinose, weakly narrowed, slightly higher than wide under antennae; occiput concave, black, grey pollinose; pair of long ocellar, pair of long vertical, and pair of shorter postvertical bristles; postocular setae relatively short, uniserial, black above, whitish below; lower postcranium with rather long white setae; eyes with microscopic white hairs. Antennae inserted in middle of head, black, longer than height of head; scape long, bare; pedicel covered with dorsal and ventral setulae; postpedicel subtriangular, with rounded apex, as long as high, covered with short hairs; arista-like stylus mid-dorsal, with microscopic hairs; length ratio of scape to pedicel to postpedicel to stylus, 9/4/6/26. Palpus short, yellow, with several hairs and 2 black apical setae; proboscis short, black, with short black hairs.
Thorax: mesonotum metallic dark-green; pleura black-green, weakly pollinose; four pairs of dorsocentral bristles; acrostichals biserial, well developed, 4 or 5 pairs; scutellum with two long strong setae and two short lateral hairs.
Legs: coxae and femora black except yellow apices, tibiae yellow, fore and mid tarsi black except most part of basitarsus yellow, hind tibia dark at apex, hind tarsus entirely black; fore coxa anteriorly with black hairs and setae of various length; mid coxa with black setae anteriorly and apically; mid and hind coxae with black external seta; fore femur with short setulae; mid and hind femora with 3 to 5 fine ventral hairs, 2/3 as long as femora diameter and 2 or 3 fine subapical anterior setae; fore tibia without conspicuous setae; mid tibia with 2 long anterodorsal, 2 short posterodorsal bristles; hind tibia with 1 or 2 anterodorsal, 3-4 posterodorsal bristles; all tibiae with apical setae; tarsi simple, with short claws and small pulvilli; coxa, femur, tibia and tarsomere (from first to fifth) length ratio: fore leg: 25/38/39/20/9/7/5/5, mid leg: 18/40/47/25/10/8/5/5, hind leg: 12/45/56/15/14/9/6/-.
Fig. 1. Asyndetus pseudoseparatus sp. nov., hypopygium, lateral aspect.
Wing: hyaline, veins brown; R1 short, ending at basal 2/5 of wing; ratio of costal section between R2+3 and R4+5 to that between R4+5 and M1+2, 14/23. R2+3 and R4+5 straight; M1+2 distinctly interrupted with broken sections partly running parallel to each other; cross-vein dm-cu absent; anal vein well developed; anal angle obtuse; lower calypter yellow, with fine white cilia; halter yellow.
Abdomen: bronze-black, with black hairs and marginal setae; sterna 4-5 developed, setose; tergum 6 glabrous; sternum 6 and segment 7 reduced; segment 8 large, rounded, with four strong black bristles; hypopygium (Fig. 1) black, small, partly concealed; epandrium flattened laterally, with left lateral foramen; hypandrium fused with epandrium, simple, short; phallus long and thin, simple; a pair of long and broad symmetrical epandrial lobes originating near base of hypandrium, with 2 apical and 1 basal setae as figured; surstylus bilobate, more or less straight; ventral lobe of surstylus gradually narrowed towards apex, bearing some short setulae and one strong middorsal seta; dorsal lobe narrow, more than half as long as ventral lobe, bearing short apical spine; postgonite narrow, slightly curved ventrally, reaching middle of surstylus; cercus small, rounded, bearing short setae.
Female: similar to male except lacking MSSC.
Measurements (mm): Body length 1.7, antenna length 0.5, wing length 1.7, wing width 0.6.
Asyndetus congensis sp. nov.
(Figs. 2−6)
Diagnosis. The new species is close to A. namibiensis, both differing from other Afrotropical species in the unusually ciliated hind tibia (see key below). Male of A. congensis has ventral setae on mid and hind femora about twice as long as diameter of corresponding femur.
Type material. HOLOTYPE ♂, D.R. Congo, Oriental Prov., Lieki village area at: 00°41.117'N, 24°14.362'E, 25.v–4.vi.2010, A.H. Kirk-Spriggs, sweeping, bush paths & village environs [BMSA]. PARATYPE ♀, same label.
Etymology. The species is named for the country of origin.
Description. Male (Fig. 2): Head (Fig. 3): vertex flat; upper occiput concave; frons wider than face, black, grey pollinose; face wide, parallel-sided and covered by silvery-white shining dust on black ground-colour; face under antenna 1.5 times as wide as basal height of postpedicel; one pair of strong ocellars; one strong vertical bristle on each side at eye margin; one postvertical on each side, not far from upper postocular seta; postocular setae relatively short, uniserial, black above, whitish below; lower post-cranium with rather long white setae. Antenna (Fig. 3) black; scape and pedicel normal; postpedicel subtriangular with rounded tip, nearly as long as high, bearing short hairs; arista-like stylus dorsal, with short segment 1 and long segment 2. Length ratio of scape to pedicel to postpedicel to stylus (segments 1 and 2), 8/6/8/5/58. Proboscis and palpus small, black; palpus whitish pollinose, with black setae.
2
3
Figs. 2−5. Asyndetus congensis sp. nov.: 2 − habitus, 3 – head, 4 – hind leg, 5 – wing.
Thorax: shining greenish-black, weakly pollinose; upper part of proepisternum with one black seta anteriorly; lower part of proepisternum with one strong and one weak black setae above coxa; 4 pairs of strong dorsocentral bristles of about equal length; additional short fine seta between 2nd and 3rd dorsocentrals, somewhat shifted; short acrostichals in biserial row; scutellum with one pair of strong widely spaced bristles and two small lateral setae.
Legs: mainly greenish-black, with black bristles and hairs, fore and mid knees dirty yellow, fore and mid tibiae pale brown; fore and mid coxae with some short and long setae anteriorly; hind coxa with one seta at base; fore femur with two ventral rows of fine setae, usually shorter than height of femur (MSSC); mid femur with 5-6 anteroventral and 5-6 posteroventral setae in middle, 2-3 times longer than height of femur (MSSC), with short anteroventrals and posteroventrals at apex; hind femur (Fig. 4) with 3-4 anteroventrals in middle and 3-4 posteroventrals behind middle, 1.5-2 times longer than height of femur (MSSC), with 2 elongate subapical anteroventral and 3 short subapical posteroventral setae; fore tibia with 1 fine dorsal at basal quarter, with 3-4 apicals and posteroventral row of elongate setulae (MSSC); mid tibia with 2 anterodorsals and 2 posterodorsals, of which upper anterodorsal bristle somewhat stronger, 3-4 apicals; hind tibia (Fig. 4) inconspicuously thickened in basal half (MSSC), with 2 strong dorsal and 1 strong posterodorsal bristles, with 3–4 shorter posterodorsals, with 7-8 fine posterior setae in basal two thirds, 1.5-2 times longer than diameter of tibia (MSSC), with double antero- and posterodorsal row of fine setae in distal two thirds, 1.5-2 times longer than diameter of tibia (MSSC), with 4 apicals; fore basitarsus with posteroventral row of elongate setulae, slightly longer than diameter of segment (MSSC); all tarsi without claws, with enlarged pulvilli (MSSC); femur, tibia and tarsomere (from first to fifth) length ratio: fore leg: 77/72/34/11/9/7/10, mid leg: 77/82/40/19/12/9/10, hind leg: 89/94/30/23/15/9/10.
Wing (Fig. 5): colorless and transparent, with brown veins; ratio of costal section between R2+3 and R4+5 to that between R4+5 and M1+2, 23/34; R4+5 and M1+2 subparallel in middle part and slightly divergent in apical part of wing; M1+2 with bend in middle of apical part, strongly weakened at bend and somewhat weakened in apical part; section of M1+2 between posterior cross-vein (dm-cu) and bend slightly longer than that between bend and wing margin (69/66); dm-cu located at level of R1; ratio of apical portion of CuA1 to dm-cu, 73/13; anal vein distinct, anal lobe developed, anal angle obtuse; calypter yellow, with simple yellow cilia; halter yellow.
Abdomen: black, with black setation; sterna 4-5 developed, setose; tergum 6 glabrous; sternum 6 and segment 7 reduced; segment 8 large, rounded, with four strong black bristles; hypopygium (Fig. 6) black, small, partly concealed; epandrium flattened laterally, with left lateral foramen; hypandrium fused with epandrium, simple, short, triangular (ventral aspect); phallus long and thin, simple; a pair of long and narrow symmetrical epandrial lobes originating near base of hypandrium, pointed at apex, with 3-4 setae as figured; surstylus bilobate, more or less straight; ventral lobe of surstylus subtriangular in basal half, narrow in distal half, with subapical ventral notch, bearing some short setulae and one strong but short middorsal seta; dorsal lobe narrow, half as long as ventral lobe, bearing short apical seta; postgonite reduced, concealed; cercus
black, small, rounded, bearing short setae.Fig. 6. Asyndetus congensis sp. nov., hypopygium, lateral aspect.
Female: similar to male except lacking MSSC, otherwise as noted: face white pollinose; hind tibia with 1 anterodorsal and 2-3 posterodorsal bristles; abdomen with anal plate projecting, rounded; hemitergites of last segment each with 4 thick spines.
Measurements (mm): Body length 2.1, antenna length 0.76, wing length 2.2, wing width 0.8.
Asyndetus savannensis sp. nov.
(Figs. 7−9)
Diagnosis. The new species is close to A. congensis and A. madagascarensis, differing in distinctly interrupted M1+2, with broken sections usually partly running parallel to each other; in acicular in distal half epandrial lobe, bearing one strong apical seta. A. congensis and A. madagascarensis have more or less weakened bend in middle of apical part M1+2; epandrial lobe bearing two apical setae in A. madagascarensis; male of A. congensis has ventral setae on mid and hind femora about twice as long as diameter of corresponding femur (see key below).
Type material. HOLOTYPE ♂, [D.R. Congo:], Congo Belge, P.N.G., Miss. H. De Saeger, II/fd/17, 29.VII.1952, H. De Saeger, 3844 [RMCA]. PARATYPES 9♂, same label; 1♂, [D.R. Congo:], Congo Belge, P.N.G., Miss. H. De Saeger, II/gd/4, 8.VIII.1952, H. De Saeger, 3923 [RMCA]; 1♂, [D.R. Congo:], Congo Belge, P.N.G., Miss. H. De Saeger, II/gd/6, 2.IX.1952, H. De Saeger, 4023 [RMCA]; 1♂, [D.R. Congo:], Congo Belge, P.N.G., Miss. H. De Saeger, II/gc/8, 9.IX.1952, H. De Saeger, 4042 [RMCA]; 1♂, [D.R. Congo:], Congo Belge, P.N.G., Miss. H. De Saeger, II/gd/4, 31.VII.1952, H. De Saeger, 3859 [RMCA]; 1♂, [D.R. Congo:], Congo Belge, P.N.G., Miss. H. De Saeger, II/gd/4, 25.VIII.1952, H. De Saeger, 3978 [RMCA]; 2♂, Senegal: M’Bour, St. ORSTOM, 7.XII.1979, D. Pluot [MNHN].
Etymology. The species is named for the ecoregion uniting two rather distant collection areas in NE Congo and Senegal, where the new species was taken.
Description. Similar to A. congensis in all respects, except for the following features. Male: Head: antenna (Fig. 7) black; length ratio of scape to pedicel to postpedicel to stylus (segments 1 and 2), 10/6/10/5/61.
Thorax: no additional short seta between 2nd and 3rd strong dorsocentrals.
7
Figs. 7−8. Asyndetus savannensis sp. nov.: 7 – antenna, 8 – wing.
Legs: mainly greenish-black, knees dirty yellow, fore and mid tibiae and bases of tarsi pale brown, hind tibia brown; all femora with two ventral rows of fine setae, half as long as height of femur (MSSC); hind tibia simple, without rows of remarkable setae; femur, tibia and tarsomere (from first to fifth) length ratio: fore leg: 86/82/40/17/12/10/11, mid leg: 92/96/47/22/15/10/10, hind leg: 95/105/34/26/15/10/13.
Wing (Fig. 8): colorless and transparent, with brown veins; ratio of costal section between R2+3 and R4+5 to that between R4+5 and M1+2, 28/46; R4+5 distinctly convex anteriorly at wing apex; M1+2 distinctly interrupted, with broken sections usually partly running parallel to each other; section of M1+2 between posterior cross-vein (dm-cu) and bend distinctly longer than that between bend and wing margin (88/72); dm-cu located right before level of R1; ratio of apical portion of CuA1 to dm-cu, 93/18.
Abdomen: black, with black setation; sterna 4–5 developed, setose; tergum 6 glabrous; sternum 6 and segment 7 reduced; segment 8 large, rounded, with four strong black bristles; hypopygium (Fig. 9) black, small, partly concealed; epandrium flattened laterally, with left lateral foramen; hypandrium fused with epandrium, simple, short, triangular (ventral aspect), with apical brush of setae; phallus long and thin, practically simple, with indistinct dorsal serration at base of swollen part; a pair of long symmetrical epandrial lobes originating near base of hypandrium, broad at base, acicular at apex (with somewhat variable length and width, in holotype slightly longer, than in some paratypes), with 1 apical and 2 subapical setae as figured (variable in location and length in paratypes); surstylus bilobate, more or less straight; ventral lobe of surstylus subtriangular in basal half, narrow in distal half, with subapical ventral notch, bearing some short setulae (variable in number and length in paratypes) and one short, but strong apicoventral spine, having no middorsal seta; dorsal lobe narrow, about half as wide as ventral lobe in middle, bearing short apical spine; postgonite bilobed, with both lobes narrow, curved ventrally, reaching apex of dorsal lobe of surstylus; cercus black,
small, rounded, bearing short setae.Fig. 9. Asyndetus savannensis sp. nov., hypopygium, lateral aspect.
Female: unknown.
Measurements (mm): Body length 2.6–2.8, antenna length 0.85, wing length 2.6, wing width 0.9.
Asyndetus namibiensis sp. nov.
(Figs. 10−12)
Diagnosis. The new species is close to A. congensis, both differing from other Afrotropical species in the unusually ciliated hind tibia (see key below). Male of A. namibiensis differs in ventral setae on mid and hind femora about half as long as diameter of corresponding femur.
Type material. HOLOTYPE ♂ [in glycerol in vial, mounted on pin], Namibia: Katima Mulilo Dist., mopane in Salambala, M4, S17°42'55'' E24°32'47'', 24-26.II.2001,
E. Marais & A.H. Kirk-Spriggs, yellow pans [BMSA]. PARATYPES 4♂, 4♀ [in alco
hol in vial, 1♂, 1♀ dried and mounted on pins], same label [BMSA].
Etymology. The species is named for the country of origin.
Description. Similar to A. congensis in all respects, except for the following fea
tures. (All specimens are quite discolourated due to long-term storage in alcohol, with major bristles brown to brown-black.). Male: Head: antenna (Fig. 10) with postpedicel trapezoidal, with rounded tip, 1.3 times as long (along ventral margin) as high at base, bearing short hairs; arista-like stylus basodorsal; palpus discolourated, but apparently dark, brownish in dried specimen. Length ratio of scape to pedicel to postpedicel to stylus (segments 1 and 2), 12/11/20/8/39.
12
Figs. 10−12. Asyndetus namibiensis sp. nov., 10 – an
tenna, 11 – hind leg, 12 – hypopygium, lateral aspect.
Thorax: upper part of proepisternum with one seta; lower part of proepisternum with one strong and one weak setae above coxa; 4 pairs of strong dorsocentral bristles of about equal length; no additional seta between 2nd and 3rd dorsocentrals; short acrostichals in biserial row; scutellum with one pair of strong bristles and two small lateral setae.
Legs: discolourated, with coxae and femora distinctly darker than tibiae; femora without strong bristles, with short anteroventrals and posteroventrals at apex; hind femur (Fig. 11) with one strong erect ventral seta at basal third, not longer than height of femur (MSSC); fore tibia with 3-4 apicals; mid tibia with 2 anterodorsals and 1-2 posterodorsals, of which upper anterodorsal bristle somewhat stronger, 3-4 apicals; hind tibia (Fig. 11) with 1-2 posterodorsals, with one dorsal and one ventral rows of elongate semi-erect setae in distal half, of which ventral setae about as long as diameter of tibia (MSSC), with 4 apicals; all tarsi without claws, with enlarged pulvilli (MSSC); femur, tibia and tarsomere (from first to fifth) length ratio: fore leg: 80/76/43/18/13/8/10, mid leg: 90/96/51/23/17/10/9, hind leg: 90/104/34/27/20/13/10.
Wing: similar to that in A. congensis; ratio of costal section between R2+3 and R4+5 to that of costal section between R4+5 and M1+2, 26/39; section of M1+2 between dm-cu and bend distinctly shorter than that between bend and wing margin (55/78); ratio of apical portion of CuA1 to dm-cu, 74/14.
Abdomen: tergum 6 almost glabrous, with at most 2 middorsal setae; segment 7 reduced; segment 8 with four strong bristles and short hairs; the bristles nearly as long as epandrium; hypopygium (Fig. 12) brown, small, partly concealed; phallus long and thin, simple; epandrial lobe subtriangular, pointed at apex, with three setae as figured; ventral lobe of surstylus elongate-triangular (dorso-lateral aspect), pointed at apex, bearing some short setulae and one strong middorsal seta; dorsal lobe narrow, rod-like, bearing weak apical seta; postgonite narrow, curved ventrally, reaching middle of surstylus; cercus small, rounded, bearing short setae.
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Female: similar to male except lacking MSSC, otherwise as noted: postpedicel subtriangular with rounded tip, nearly as long as high; mid tibia with both anterodorsals equally strong; hind tibia with few short dorsal setae.
Measurements (mm): Body length 2.2 (dry) – 2.7 (in alcohol), antenna length 0.7, wing length 2.2, wing width 0.9.
Asyndetus madagascarensis sp. nov.
(Figs. 13−15)
Diagnosis. The new species is close to A. virgatus, differing in longer postpedicel, nearly 2 times longer than high, and presence of the large suboval epandrial lobe (see key below).
Type material. HOLOTYPE ♂ [in glycerol in vial, mounted on pin], MADAGASCAR: Fia[narantsoa Province], Ranomafana [National Park, 21°00'S 47°30'E], 19.I.1992, A. Pauly, forêt (IRSNB). PARATYPES 14♂, 7♀ [in alcohol in vial, 1♂, 1♀ dried and mounted on pins], same label [IRSNB].
Etymology. The species is named for the country of origin.
Description. Similar to A. congensis in all respects, except for the following features. (All specimens are quite discolourated due to long-term storage in alcohol, with major bristles brown to brown-black.). Male: Head: antenna (Fig. 13) with postpedicel large, trapezoidal, with rounded tip, 1.8 times as long (along ventral margin) as high at base, bearing short hairs; arista-like stylus basodorsal; palpus discolourated, but apparently dark, brownish in dried specimen. Length ratio of scape to pedicel to postpedicel to stylus (segments 1 and 2), 12/10/22/6/36.
13
14
Figs. 13−14. Asyndetus madagascarensis sp. nov.: 13 – antenna, 14 – wing.
Thorax: upper part of proepisternum with one seta; lower part of proepisternum with one strong and one weak setae above coxa; 5 pairs of strong dorsocentral bristles; short acrostichals in biserial row; scutellum with one pair of strong bristles and two small lateral setae.
Legs: discolourated, with coxae and femora distinctly darker than tibiae; all femora with double row of long ventral setae along entire length, at least half as long as height of femur, in addition to subapical ventral bristles (MSSC); at apices those setae nearly as long as height of corresponding femur; fore tibia with 3-4 apicals; mid tibia with 2 anterodorsals and 2 posterodorsals, of which upper anterodorsal bristle somewhat stronger, 3-4 apicals; hind tibia with one dorsal row of about 8 short setae, of which one middorsal seta strongest, slightly longer than diameter of tibia (MSSC), with 4 apicals; all tarsi without claws, with enlarged pulvilli (MSSC); femur, tibia and tarsomere (from first to fifth) length ratio: fore leg: 75/71/37/14/10/6/9, mid leg: 84/80/42/17/11/8/8, hind leg: 88/98/27/22/14/9/8.
Wing (Fig. 14): similar to that in A. congensis; ratio of costal section between R2+3 and R4+5 to that of costal section between R4+5 and M1+2, 27/37; wing vein dm-cu faint, located before level of R1; section of M1+2 between dm-cu and bend distinctly longer than that between bend and wing margin (82/63); ratio of apical portion of CuA1 to dm
40
cu, 89/9.
Abdomen: tergum 6 glabrous; segment 7 reduced; segment 8 with four strong bristles and short hairs; hypopygium (Fig. 15) small, partly concealed; phallus long and thin, simple; epandrial lobe suboval, with three long setae as figured; ventral lobe of surstylus elongate-triangular (dorso-lateral aspect), acute at apex, bearing some short setulae and one or two strong middorsal setae; dorsal lobe narrow, rod-like, bearing short apical spine; postgonite narrow, curved ventrally, reaching middle of surstylus;
cercus small, rounded, bearing moderately long setae.Fig. 15. Asyndetus madagascarensis sp. nov. hypopygium, lateral aspect.
Female: similar to male except lacking MSSC, otherwise as noted: postpedicel subtriangular with acute tip, as long as high; mid tibia with both anterodorsals equally strong; hind tibia with 2 anterodorsal and about 5 short dorsal setae.
Measurements (mm): Body length 2.1 (dry) – 2.6 (in alcohol), antenna length 0.65, wing length 2.1, wing width 0.85.
Doubtful species
Asyndetus indifferens Curran, 1926b: 411. Remarks: A female holotype (South African Museum, Cape Town, not seen) was incompletely described, being compared with a male of A. crassitarsis Curran, 1926b, differing from the latter in brown fore tibia, the character being common in females of
A. virgatus (Curran, 1926a) and A. decaryi (Parent, 1929b). I think the A. indifferens may be conspecific to either A. virgatus or A. decaryi. Distribution: Type locality: South Africa: Zululand, M’fongosi. Afrotropical: South Africa. Asyndetus inermis Parent, 1937: 7. Type material examined: ♀ [holo]type: [DR Congo:] Congo-belge, Rives Busira, VI.1936, J. Ghesquière / R. Mus. Hist. Nat. Belg. 10482 / Asyndetus inermis n.sp. ♀, O. Parent / O. Parent det. 1936: Asyndetus inermis n.sp. / tête disparue du course duretour du determinateur [RMCA]. Remarks: The holotype examined is strongly damaged. According to the original description, the species is close to the A. albifrons female, differing in reduced number of bristles on tibiae (1 antero-, 2 posterodorsals on mid tibia and 2 antero-, 2 posterodorsals on hind tibia in A. inermis vs. 2 antero-, 2 posterodorsals on mid tibia and 2 antero-, 4 posterodorsals on hind tibia in A. albifrons and all Afrotropical species of the genus)
and in distinctly interrupted wing vein M1+2 (weakly interrupted in distal part in A. albifrons). Legs black except fore and mid knees. Distribution: Type locality: Congo-belge, Rives Busira. Afrotropical: DR Congo.
Key to Afrotropical species of Asyndetus based on male characters
Remarks: Females usually cannot be identified without males of the same series; A. inermis and A. indifferens, known only from females, are not included.
1. At least anterior four tarsi with claws, with small pulvilli; M1+2 distinctly interrupted with broken sections partly running parallel to each other; wing vein dm-cu indistinct; 1.7–2.2 mm .................................................................................. pseudoseparatus sp. n.
– At least fore tarsus without claws, with enlarged pulvilli; M1+2 usually with bend in middle of apical part, more or less weakened at bend; wing vein dm-cu usually distinct .. ..........................................................................................................................................2
2. Fore basitarsus slender; segments 4–5 of fore tarsus dilated and flattened; 3 mm ......... ............................................................................................................. crassitarsis Curran
– Fore tarsus unmodified .................................................................................................3
3. Hind tarsus with claws, with small pulvilli; 2.5 mm .......................... albifacies Parent
– All tarsi without claws, with enlarged pulvilli ..............................................................4
4. Hind tibia with ventral row of semi-erect setae, at least as long as diameter of tibia ...5
– Hind tibia without remarkable ventral row of long setae ..............................................7
5. Hind tibia with regular or irregular row of elongate ventral or anteroventral setulae along entire length, not longer than diameter of tibia; epandrial lobe absent; 2.5–3 mm ... ............................................................................................... virgatus Curran (see below)
– Hind tibia with regular ventral row of semi-erect setae in middle or in distal half, at least as long as diameter of tibia and distinctly longer than other setulae in the same and other rows; epandrial lobe well developed ......................................................................6
6. Mid and hind femora with double rows of erect ventral setae, about 2 times longer than height of femur; epandrial lobe long and narrow; 2.1 mm ...............congensis sp. n.
– Mid femur with ventral setae, at most as long as height of femur; hind femur ventrally almost bare, with one strong erect ventral seta at basal third, not longer than height of femur; epandrial lobe large, triangular; 2.2 mm ...................................namibiensis sp. n.
7. Mid and hind femora with double row of ventral setae along entire length, about half as long as diameter of femur, in addition to subapical ventral bristles; postgonite long and narrow .......................................................................................................................8
– Mid and hind femora with only subapical ventral bristles; postgonite usually short, concealed .......................................................................................................................10
8. M1+2 distinctly interrupted, with broken sections usually partly running parallel to each other; epandrial lobe pointed in distal half, long and narrow, bearing one strong apical seta ............................................................................................. savannensis sp. n.
– M1+2 with bend in middle of apical part, more or less weakened at bend; epandrial lobe either absent or bearing two apical setae ..................................................................9
9. Postpedicel 1.5–2 times longer than high; wing vein dm-cu faint; epandrial lobe large, suboval, with three long setae; surstylus with dorsal lobe much narrower than ventral lobe; 2.1 mm ................................................................................ madagascarensis sp. n.
– Postpedicel at most slightly longer than high; wing vein dm-cu strong; epandrial lobe absent; surstylus with dorsal lobe wide, nearly as wide as ventral lobe at apex; 2.5–3 mm ............................................................................................................virgatus Curran 10. Halter brown; M1+2 distinctly interrupted; hind tibia with 1 antero- and 1 posterodorsal bristles at base and one row of dorsal setae in distal half ............... amaphinius Séguy
– Halter clear yellow; M1+2 usually with bend in middle of apical part, more or less weakened at bend; hind tibia differently setose .............................................................11
11. Section of M1+2 between posterior cross-vein (dm-cu) and bend about as long as that between bend and wing margin; epandrial lobe short and wide, distinctly shorter than surstylus, bearing two apical setae; 2–3 mm ........................................... latifrons (Loew)
– Section of M1+2 between posterior cross-vein (dm-cu) and bend distinctly longer than that between bend and wing margin ..............................................................................12
12. Frons silvery-white pollinose; fore femur with double ventral row of long setae; 2.5 mm ........................................................................................................... albifrons Parent
– Frons grey pollinose; fore femur with short setae; epandrial lobe long and slender, rod-like, bearing two apical setae; 2 mm ....................................................decaryi Parent
Appendix. Notes on some Palaearctic and Oriental species General remarks: As noted above, the presence or absence of acrostichal setae is the only appropriate character for distinguishing Asyndetus from Cryptophleps (Capellari & Grichanov, 2012). Therefore, at least three Palaearctic species described in the Asyndetus (A. izius Negrobov, 1973, A. minutus Negrobov et Shamshev, 1986 and A. vividus Negrobov et Shamshev, 1986) may be recombined with Cryptophleps. Nevertheless, my investigation of the hypopygial morphology of species of the two genera (see figures in this and cited papers) assumes that the genera will be synonymised in future.
Asyndetus albipalpus Loew, 1871: 295; Negrobov, 1973: 158. Type material examined (here designated): Lectotype: ♂, [Uzbekistan: upper stream of Syr Darya River:] “Сыръ Дарья” [ZIN]. Paralectotypes: 2♂, same locality [ZIN].
Remarks: Describing new species from the A.P. Fedtshenko’s Turkistan expedition (1868-1871), H. Loew everywhere mentioned “Sarawschan Thal” (Zeravshan River Valley) as the type locality of his newly described species. Nevertheless, A.P. Fedtshenko collected the material from a much larger territory of the former Khanate of Kokand, from Kattaqurghon and Samarqand cities in the West to Fergana Valley and Alai Mountains in the East (e.g., Negrobov, 1981) within the present countries Kyrgyzstan, Tajikistan and Uzbekistan. Some collection sites were also located in the present Kazakhstan. Most part of the identified material was returned to ZMU at the end of XIX century, while some voucher specimens (including Loew’s syntypes) were retained in the Natural History Museum, Berlin, Germany (MFN; Negrobov, 1981). A part of the ZMU material collected by Fedtshenko was probably borrowed or taken in exchange by
A.A. Stackelberg (ZIN) before the WWII and was recently found in ZIN collection. It contains apparently mixture of identified and unidentified specimens under original labels, comprising locality number (separately) and brief description of a locality (in Cyrillic letters). Six specimens bear old hand-written museum labels with species names (including two specimens of Loew’s species with abbreviation “n. sp.”; one of the names is unpublished), followed by a more or less long row of unlabeled specimens. Some specimens bear hand-written (by Stackelberg) identification labels including types (with abbreviation “n. sp.”) of at least two species, i.e. the holotype of Syntormon turanicus Stackelberg, 1927 and two syntypes of Teuchophorus rohdendorfi Stackelberg, 1927, in addition to possible syntypes (with only locality labels) of Fedtshenkomyia chrysotymoides Stackelberg, 1927 and Rhaphium turanicola (Stackelberg, 1927).
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Regarding Asyndetus, the ZIN old material examined comprises a series of specimens followed by a male (with broken abdomen) bearing an old hand-written museum label “albipalpus Lw.” and Stackelberg’s hand-written label “Asyndetus albipalpis Lw.”. The series contains three Asyndetus species, of which three males (with a locality label “Syr Darya” in Russian) fully correspond to the original Loew’s description of A. albipalpus (Loew, 1871) and generally correspond to a brief description of A. albipalpus by Negrobov (1973), which was based on Mongolian material and may belong to a different species. The other specimens belong to A. longicornis Negrobov, 1973 and A. mixtus Negrobov et Shamshev, 1986 (see below). Therefore, lectotype and paralectotypes are here designated to fix the current taxonomic concept of A. albipalpus and ensure consistent future interpretation.
Distribution: Type locality: Uzbekistan: upper stream of Syr Darya River (originally published as “Turkestan, Sarawschan Thal”). Palaearctic: ?Mongolia, Tajikistan, Uzbekistan.
Asyndetus barbiventris Stackelberg, 1952: 402; Negrobov, 1973: 159. Material examined: 1♂, Tajikistan: Khatlon Prov., Farkhor distr., Novobod, 37.551°N, 69.460°E, 475 m asl, 6.VI.2010, water, pastures, K. Tomkovich [ZMU]; 1♂, [Kazakhstan or Uzbekistan]: Bairakum / 4 [pink label; ex coll. A.P. Fedtshenko 1871; ZIN]; Distribution: Type locality: Tajikistan: Tigrovaya balka, river Pyandzh, Kirovobad. Palaearctic: ?Kazakhstan, Tajikistan, ?Uzbekistan. First record after description.
Asyndetus chaetifemoratus Parent, 1925: 162. Material examined: 3♂, 1♀, Israel: Mt. Hermon, 1600 m, 5.IX.1981, A. Freidberg [TAU]. Distribution: Type locality: Egypt: Baharia Oasis. Palaearctic: Egypt, Israel. New for Israel. First record after description.
Asyndetus longicornis Negrobov, 1973: 160. Material examined: 1♂, 1♀, [Uzbekistan: upper stream of Syr Darya River:] “Сыръ Дарья” [ZIN]. 2♂, 1♀, [Kazakhstan: Kosaral:] “Косаралъ” /24 [ZIN]; 1♂, [Tajikistan or Uzbekistan: Zeravshan Valley:] “Заравш. д.” /1 [ZIN]; 1♂, Russia: Astrakhan Region, Ikryanoe district, Zyuzino, 45.751°N, 47.678°E, 8–9.V.2010, water, pastures, K. Tomkovich [ZMU]. Additional material examined: 1♀, [Tajikistan or Uzbekistan: Zeravshan Valley:] “Заравш. д.” /6 [ZIN]; 2♀, [Uzbekistan: Keles:] “Келесъ” /22 [ZIN]; 2♀, [Uzbekistan: Karak:] “Каракъ” /7 [ZIN]; 1♀, [Kazakhstan:] Taldyqorghan Region, Katutau Mts., Kuibynskoe Gorge, 25 km NWN Koktal, 30.VI.1988, Tanasiichuk [ZIN]; 1♀, [Turkmenistan:] 30 km E Nebit-Dag, S slope of Bolshoi Balkhan Ridge, 9.V.1984, Tanasiichuk [ZIN]. Remarks: A. longicornis males examined show some extent of variation in shape of antennal postpedicel (with ovate to subtriangular apex) and setation of phallus (aedeagus); the characters were considered diagnostic by Pârvu (1989) when he described his new species A. negrobovi from Romania. E.g., two males collected in Kosaral differ from each other in subapical setation of phallus (in addition to shape of postpedicel): weaker (as shown by Negrobov, 1973: Fig. 24) or stronger dorsal setation with two basal dents long and narrow (as shown by Pârvu, 1989: Fig. 5). Therefore, A. longicornis can repre
44
sent a complex of widely distributed sibling species (including A. negrobovi) or different phenotypes of the same species. Distribution: Type locality: “Mongolei, Südgobi aimak, 40 km SSO von Nomagon, Salzbodenwiese, in der Nähe von der Wasserquelle”. Palaearctic: China (Inner Mongolia), Hungary, Kazakhstan, Mongolia, ?Romania, Russia (Astrakhan), Tajikistan, Turkmenistan, Uzbekistan. New for Kazakhstan, Russia, Tajikistan, Turkmenistan and Uzbekistan.
Asyndetus mixtus Negrobov et Shamshev, 1986: 48. Material examined: 1♂ (with broken abdomen), [Tajikistan or Uzbekistan: Zeravshan Valley:] “Заравш. д.” /4. / “albipalpus Lw.” [old hand-written museum label] / “Asyndetus albipalpis Lw., Stackelberg det.” (ZIN); 1♂, [Tajikistan or Uzbekistan: Zeravshan Valley:] “Заравш. д.” /28 (ZIN); 1♂, [Tajikistan: Vorukh:] “Ворухъ / 20” (ZIN). Remarks: Negrobov & Shamshev (1986) referred the type locality of the species in error to Turkmenistan instead of Uzbekistan. Distribution: Type locality: [Uzbekistan]: Samarkand, Kattakurgan district, Kumak. Palaearctic: Tajikistan, Uzbekistan. New for Tajikistan.
Asyndetus separatus (Becker), 1902: 56 (Meringopherusa); Becker, 1918: 78 (Asyndetus); Negrobov, 1973: 161. Asyndetus lateinterruptus Strobl, in Czerny, Strobl, 1909: 190 (as late-interruptus). Type locality: Italy and Austria: “Monfalcone bei Triest; Admont” (synonymised by Becker, 1918: 78) Material examined: 1♂, [Egypt:] Fayūm, III, 44741 / Meringopherusa separata Beck., det. Becker [ZIN]; 1♀, [Egypt:] Kairo, XI, 44277 [ZIN]; 3♂: Cyprus, Famagusta, 912.VII.1939 (Håkan Lindb.) [ZMH]; 3♀, Israel: Ein Hajla, 11.V.1977, A. Freidberg [TAU]. Remarks: A male from Fayūm (Egypt) examined may belong to the type series of A. separatus. It corresponds to the description and figure provided by Becker (1902). The male genitalia preparation was made by the author of this paper, showing its strong similarity with the A. pseudoseparatus described in this paper (see Fig. 1), but differing from the figure provided by Negrobov (1973) in shape of epandrial lobe and in simple phallus (without setation). In addition, a male from Fayūm has two pedunculate apical setae on the left epandrial lobe and three apical setae (of which one seta is pedunculate) on the right epandrial lobe. Negrobov (1973) studied the material collected from Tajikistan, and his pictures may belong to another species. Distribution: Type locality: Egypt: Alexandrien, Fayūm. Palaearctic: Algeria, Cyprus, Egypt, Greece, Iraq, Israel, Libya, Spain, Tunisia, ?Tajikistan. New for Israel.
Asyndetus thaicus Grootaert et Meuffels, 2002: 42; Wang et al., 2007: 151. Material examined: 2♂, India: Gujarat, Naliya, env. Kothara, riv. Nayera, 23.137°N, 68.931°E, 5.X.2012, K. Tomkovich [ZMU]; 1♂, 2♀, India: Goa, Palolem, ~15.018°N, 74.018°E, 3-9.II.2009, K. Tomkovich [ZMU]. Remarks: This is the second Asyndetus species found in India, which is still undercollected. The wide areas of A. latifrons and A. thaicus distribution assume the presence of more Oriental, Palaearctic and Afrotropical species of the genus in this country. Distribution: Type locality: Thailand: Rayong Province, Ko Samed, Tarn Tawan. Oriental: China (Yunnan), India (Goa, Gujarat), Thailand. New for India.
Acknowledgements
I am sincerely grateful to Dr. E.P. Nartshuk and L.A. Kuznetsova (St. Petersburg), Drs. Nikita Vikhrev and Andrei Ozerov (Moscow), Drs. Patrick Grootaert and Marc De Meyer (Brussels), late Dr. Loïs Matile (Paris), Dr. Ashley H. Kirk-Spriggs (Bloemfontein), Dr. Mike Mostovski (Pietermaritzburg), Roy Danielsson (Lund), Dr. Amnon Freidberg (Tel Aviv), Dr. C.F. Kassebeer (Kiel), Dr. Thomas Pape (Copenhagen), Dr. Laszlo Papp (Budapest), Dr. Brian Pitkin (London), and Dr. Pekka Vilkamaa (Helsinki) for their kindness in furnishing an opportunity to study the collections of their museums. Renato Capellari (São Paulo) has made some useful comments. This paper was partly supported by the grant of the Russian Foundation for Basic Research N 11-04-01051-a to Oleg P. Negrobov.
References
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CHECK-LIST OF PREDATORY FLIES OF THE FAMILY DOLICHOPODIDAE (DIPTERA) IN THE FAUNA OF RUSSIA
O.P. Negrobov1, O.V. Selivanova1, O.O. Maslova2, M.A. Chursina1
1Voronezh State University, Voronezh, 394006, Russia. E-mail: negrobov@list.ru 2Voronezh State Pedagigical University, Voronezh, 394043, Russia
О.П.Негробов, О.В.Селиванова, О.О.Маслова, М.А.Чурсина. Справочный список хищных мух-зеленушек семейства Dolichopodidae (Diptera) фауны России
Приведен обзор изученности фауны семейства Dolichopodidae (Diptera) России. Первые сведения по Dolichopodidae России с описанием ряда новых видов включены в работы зарубежных и российских авторов XIX-XX веков (Dwigubsky, 1802; Fischer von Waldheim, 1819; Gimmerthal, 1847; Eversmann, 1834; Остен-Сакен, 1858; Motschulsky, 1859; Loew, 1848, 1871; Kowarz, 1877; Порчинский, 1874; А. Федченко, 1868; Б. Федченко, 1891, 1892; Becker, 1900, 1915, 1923; Lundström, Frey 1913; Frey, 1915, 1918; Wnukowsky, 1932, 1936; Parent, 1925, 1927, 1930). А.А. Штакельбергом (1919–1971) было впервые для России указано 68 видов долихоподид, а также описано много новых видов с юга Дальнего Востока, из Сибири и других регионов нашей страны. Е.С. Смирновым (1948а, 1948б) было описано 18 видов рода Dolichopus из Приморья и 2 вида рода Hercostomus (Смирнов, Негробов, 1977, 1979). К началу 1960-х гг. наиболее полно была исследована территория Ленинградской области, для которой было отмечено 205 видов Dolichopodidae (Штакельберг, 1962). О.П. Негробов с соавторами (1963–2012) описал из разичных регионов России более 200 новых для науки видов. И.Я. Гричановым (1979-2012) получены новые фаунистические данные и описан ряд новых видов, в том числе из европейской части России, Сибири и Дальнего Востока. Работы Б.И Вольфова и С.Ю. Кустова (2006-2010) посвящены фауне северо-западной части Кавказа. В настоящей работе впервые публикуется список хищных мух-зеленушек Dolichopodidae фауныРоссии, включающий 52 рода, 735 видов и подвидов.
Воронежский государственный университет, Воронежский государственный педагогический университет, Воронеж.
KEY WORDS: Diptera, Dolichopodidae, Russia, check-list.
Introduction
The first data on Dolichopodidae of Russia with descriptions of some new species were published by European and Russian authors (Dwigubsky, 1802; Fischer von Waldheim, 1819; Gimmerthal, 1847; Eversmann, 1834; Osten-Saken, 1858; Motschulsky, 1859; Loew, 1848-1873; Kowarz, 1873, 1877, 1889; Porchinski, 1874; А. Fedtchenko 1868; B. Fedtchenko, 1891, 1892; Becker, 1900, 1915, 1923; Lundström, Frey, 1913; Frey, 1915, 1918, 1933; Wnukowsky, 1932, 1936; Parent, 1925, 1927, 1930).
A.A. Stackelberg (1919–1971) recorded 68 species of Dolichopodidae from Russia for the first time and described many new species from the South of the Far East, from Siberia and some other regions of the country. In 1960s the Leningrad Region was the most studied territory in Russia with 205 known species of Dolichopodidae (Stackelberg, 1962). E.S. Smirnov (1948a, 1948b) described 18 species of the genus Dolichopus from Primorye and two species of the genus Hercostomus (Smirnov, Negrobov, 1977, 1979). More than 200 species new to science were described from various territories of Russia by O.P. Negrobov and coauthors (1963–2012). New faunistic data were obtained and a number of new species were described by I.Ya. Grichanov (1979–2012) from the European part of Russia, Siberia and the Far East. Works of B.I. Volfov and S.Yu. Kustov (2006-2010) were devoted to the fauna of northwestern part the Caucasus.
48
There were also a number of publications on larvae of the genus Medetera being predatious on bark beetles, on ecology and ethology of Dolichopodidae. So far 52 genera, 735 species and subspecies are known on the territory of Russia. The number of species and subspecies in genera are as follows: Achalcus – 5, Acropsilus – 1, Anepsiomyia – 1, Aphrosylus – 1, Argyra – 25, Asyndetus – 4, Campsicnemus – 27, Chrysotimus – 6, Chrysotus – 37, Diaphorus – 16, Diostracus – 5, Dolichophorus – 1, Dolichopus – 179, Epithalassius – 1, Guzeriplia – 2, Hercostomus – 54, Hydrophorus – 32, Hypophyllus – 3, Lamprochromus – 4, Liancalus – 1, Ludovicius – 1, Machaerium – 1, Medetera – 100, Melanostolus – 3, Mesorhaga – 2, Micromorphus – 5, Nematoproctus – 4, Neurigona – 22, Nepalomyia – 1, Orthoceratium – 1, Paraclius – 1, Peloropeodes – 1, Peodes – 3, Poecilobothrus – 7, Pseudoxanthochlorus – 1, Rhaphium – 62, Scellus – 5, Sciapus – 19, Setihercostomus – 1, Sphyrotarsus – 1, Suschania – 1, Sybistroma – 8, Sympycnus – 8, Syntormon – 22, Systenus
– 4, Tachytrechus – 9, Telmaturgus – 1, Teuchophorus – 6, Thinophilus – 5, Thrypticus – 20, Vetimicrotes – 1, Xanthochlorus – 5.
Check-list
Achalcus Loew, 1857.
Achalcus cinereus (Walker, 1851) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Achalcus flavicollis (Meigen, 1824) – Leningrad reg. (Stackelberg, 1925, 1962), Achalcus melanotrichus Mik, 1878 – Pskov reg. (Grichanov, 1998), Achalcus polleti Negrobov et Selivanova, 2010 – Amur reg. (Negrobov, Selivanova, 2010), Achalcus thalhammeri Lichtwardt, 1913 – Pskov reg. (Przhiboro, Grichanov, 2003).
Acropsilus Mik, 1878.
Acropsilus niger (Loew, 1869) – Russia (Parent, 1938).
Anepsiomyia Bezzi, 1902.
Anepsiomyia flaviventris (Meigen, 1824) – Russia (Parent, 1938).
Aphrosylus Walker, 1851.
Aphrosylus ferox Haliday, 1851 – Russia (Parent, 1938).
Argyra Macquart, 1834.
Argyra argentina Meigen, 1824 – Leningrad reg. (Stackelberg, 1925), North Caucasus (Negrobov, 1967 с, Negrobov, Duhanina, 1984, Grichanov, Volfov, Kustov, 2006), Argyra argyria (Meigen, 1824) – North Caucasus (Negrobov, 1967 с), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Vologda reg. (Grichanov, 2006 b), Argyra atriceps Loew, 1857 – Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 с), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Argyra auricolis (Meigen, 1824) – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Argyra diaphana (Fabricius, 1775) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1922, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 с, 1967 а), Ryazan reg. (Negrobov, Pogonin, 1984), North Caucasus (Grichanov, Volfov, Kustov, 2006), Kursk reg. (Grichanov, 2007a), Ryazan reg. (Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Argyra elongata (Zetterstedt, 1843) – Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1967 a), Vologda reg. (Grichanov, 2006 b), Argyra flavida Negrobov, 1973 – Primorye (Negrobov, 1973 c), Argyra grata Loew, 1857 – Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1972), Argyra ilonae Gosseries, 1989 (synonym – Argyra confinis (Zetterstedt, 1849) – Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007, Grichanov, 2012 c), Adygea (Grichanov, Volfov, Kustov, 2009), Argyra leucocephala (Meigen, 1824) – Volga River basin (Eversmann, 1834), Leningrad reg. (Stackelberg, 1922, 1925, 1962), Voronezh reg. (Negrobov, 1963, Negrobov, 1967 a), North Caucasus (Negrobov, 1967 с), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Krasnodar Territory (Grichanov, 2012c), Argyra magnicornis (Zetterstedt, 1838) – Murmansk reg. (Frey¸ Lundsrom, 1913), Russia (Parent, 1938), Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1963, Negrobov, 1967 a), Krasnodar Territory (Negrobov, Duhanina, 1984), Argyra negrobovi Grichanov & Shamshev, 1993
– Khabarovsk Territory (Grichanov, Shamshev, 1993, Grichanov, 2006 a), Argyra oreada Negrobov, 1973 – North Caucasus (Negrobov, 1973 c, Grichanov, 2012 c), Adygea (Grichanov, Volfov, Kustov, 2009), Argyra pulata Negrobov et Maslova, 2003 – Ural (Negrobov, Maslova, 2003), Argyra setimana Loew, 1859 – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Murmansk reg. (Grichanov, 2004), Argyra setulipes Becker, 1918 – Orenburg reg. (Becker, 1918, Negrobov, Selivanova, 2005), Argyra shamshevi Selivanova et Negrobov 2007 – Primorye (Selivanova, Negrobov, 2007), Argyra skufjini Negrobov, 1965 – North Caucasus (Negrobov, 1965 b, Negrobov, 1967 c), Argyra spoliata Kowarz, 1878 – Russia (Parent, 1938), Leningrad reg. (Stackelberg, 1962), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Khanty-Mansi autonomous region (Grichanov, 2010 a), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Lipetsk reg., Krasnodar Territory, Sayan Mountains, Buryatia, Irkutsk reg., Krasnoyarsk Territory, Primorye , Amur reg. (Selivanova, Negrobov, Maslova, 2012), Argyra subarctica Ringdahl, 1920 – Baikal (Negrobov, 1976 b), North Caucasus (Grichanov, 1998), Adygea (Grichanov, Kustov, Volfov, 2006), Altai (Negrobov, Barkalov, 2009), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Argyra submontana Negrobov et Selivanova, 2005 – Krasnodar Territory (Negrobov, Selivanova, 2005, Grichanov, 2012 c), Argyra superba Takagi, 1960 – Kuril Islands (Negrobov, 1976 b), Argyra ussuriana Negrobov, 1973 – Primorye (Negrobov, 1973 c), Argvra venevitinovensis Selivanova et Negrobov 2007 – Voronezh reg. (Selivanova, Negrobov, 2007 a), Argyra vestita (Weidemann, 1817) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Vologda reg. (Grichanov, 2006 b), Krasnodar Territory (Grichanov, 2012 c).
Asyndetus Loew, 1869.
Asyndetus diaphoriformis Negrobov et Shamshev, 1986 – Primorye (Negrobov, Shamshev, 1986a), Asyndetus latifrons (Loew, 1857) – Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1963, Negrobov, 1968 b, Negrobov, 1972), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Adygea (Grichanov, Volfov, Kustov, 2009), Asyndetus minutus Negrobov et Shamshev, 1986 – Khabarovsk Territory (Negrobov, Shamshev, 1986 a, Grichanov, 2006 a), Asyndetus vividus Negrobov et Shamshev, 1986 – Primorye (Negrobov, Shamshev, 1986 a).
Campsicnemus Haliday, 1851.
Campsicnemus argyropterus Negrobov et Shamshev, 1985 – Yakutia (Negrobov, Shamshev, 1985), Khabarovsk Territory (Grichanov, 2006 a), Campsicnemus armatus (Zetterstedt, 1849) – Murmansk reg. (Frey, 1915), Karelia (Becker, 1918), Arkhangelsk reg., Murmansk reg., Yamal-Nenets autonomous region, Yakutia, Kamchatka (Negrobov, Zlobin, 1978), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Campsicnemus articulatellus (Zetterstedt, 1843) – Pskov reg. (Przhiboro, Grichanov, 2003), Campsicnemus bagachanovae Grichanov et Volfov, 2009 – Yakutia (Grichanov, Volfov, 2009), Campsicnemus compeditus Loew, 1857 – Siberia (Frey, 1915), Russia (Parent, 1938), Leningrad reg. (Stackelberg, 1962), Buryatia, Yakutia, Kamchatka (Negrobov, Zlobin, 1978), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Campsicnemus curvipes (Fallén, 1823) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1967a), North Caucasus (Negrobov, 1965 а, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Krasnodar Territory (Negrobov, Duhanina, 1984, Grichanov, 2006 a, 2012 c), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Kursk reg. (Grichanov, 2007a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Kaluga reg. (Grichanov, 2010b), Perm reg. (Grichanov, 2012), Campsicnemus dasycnemus Loew, 1857 – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Campsicnemus femoratus Ringdahl, 1949 – Krasnoyarsk Territory (Negrobov, Zlobin, 1978, Pogonin, Negrobov, 2008), Campsicnemus filipes Loew, 1859 – Voronezh reg. (Negrobov, 1965 b, 1965 c, 1966 а), North Caucasus (Grichanov, Volfov, Kustov, 2006), Campsicnemus konstantini Grichanov, 2011 – Astrakhan reg. (Grichanov, 2011), Campsicnemus lineatus Negrobov et Zlobin, 1978 -Primorye (Negrobov, Zlobin, 1978), Campsicnemus loripes (Haliday, 1862) – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Campsicnemus lumbatus Loew, 1857
– Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1966 а, 1967 a), North Caucasus (Negrobov, 1967d, Grichanov, Volfov, Kustov, 2006), Orenburg reg. (Negrobov, Zlobin, 1978), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Moscow reg., Voronezh reg. (Grichanov, 2012), Campsicnemus magius (Loew, 1845) – North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Yakutia (Negrobov, Chalaya, 1991), Astrakhan reg. (Grichanov, 2011), Campsicnemus marginatus Loew, 1857 – Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1965c), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Campsicnemus paradoxus (Wahlberg, 1844) – Yakutia (Negrobov, Zlobin, 1978, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Campsicnemus picticornis (Zetterstedt, 1843) – Leningrad reg. (Stackelberg, 1925, 1962), Kamchatka (Parent, 1930), Russia (Parent, 1938), Yakutia, Buryatia, Sayan Mountains, Primorye (Negrobov, Zlobin, 1978), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Jewish AR, Khabarovsk Territory (Grichanov, 2006 a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Novosibirsk reg. (Grichanov, 2012), Campsicnemus pumilio (Zetterstedt, 1843) (synonym – Campsicnemus pectinulatus Loew, 1864) – Moscow reg. (Fedtchenko A., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c), Yakutia, Krasnodar Territory (Negrobov, Zlobin, 1978), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Novgorod reg. (Grichanov, 2012), Campsicnemus pusillus (Meigen, 1824) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), Irkutsk reg., Primorye (Negrobov, Zlobin, 1978), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Vologda reg. (Grichanov, 2006 b), North Caucasus (Grichanov, Volfov, Kustov, 2007), Kabardino-Balkaria (Grichanov, Volfov, 2009), Ivanov reg. (Grichanov, 2012), Campsicnemus scambus (Fallén, 1823) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915, Negrobov, 1974), Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1965 c, 1966 а, 1967 a), North Caucasus (Negrobov, 1967 c), Altai, Khabarovsk Territory (Negrobov, Zlobin, 1978), Primorye (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984), Yakutia (Negrobov, Chalaya, 1991), Vologda reg. (Grichanov, 2006 b), Khabarovsk Territory (Grichanov, 2006a), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Khanty-Mansi autonomous region (Grichanov, 2010 a), Novgorod reg. (Grichanov, 2012), Campsicnemus simplicissimus Strobl, 1906 – Russia (Negrobov, 1977 b), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Campsicnemus umbripennis Loew, 1856 – Leningrad reg. (Stackelberg, 1925), Russia (Parent, 1938), Krasnodar Territory (Negrobov, Duhanina, 1984), North Caucasus (Grichanov, Volfov, Kustov, 2006, Grichanov, Volfov, Kustov, 2007), North Caucasus, Adygea (Grichanov, Tomkovich, 2009), Krasnodar Territory (Grichanov, 2012 c), Campsicnemis unipunctatus Negrobov et Zlobin 1978 – Primorye (Negrobov, Zlobin, 1978, Grichanov, 2012), Campsicnemus varipes Loew, 1859 – Krasnodar Territory (Negrobov, 1965 b, Negrobov, Duhanina, 1984), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Yakutia (Grichanov, Volfov, 2009), Campsicnemis versicoloris Negrobov et Zlobin, 1978 – Primorye (Negrobov, Zlobin, 1978), Campsicnemis vtorovi Negrobov et Zlobin 1978 – Kirgistan (Negrobov et Zlobin 1978), Voronezh reg. (Grichanov, 2012), Campsicnemus zlobini Grichanov 2012 – Primorye (Grichanov, 2012a).
Chrysotimus Loew, 1857.
Chrysotimus concinnus (Zetterstedt, 1845) – Russia (Parent, 1938), Chrysotimus flavisetus Negrobov 1978 – Primorye (Negrobov, 1978b), Chrysotimus molliculus (Fallén, 1823) – Moscow reg. (Fedtchenko A., 1868), Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a, 1967 c), Ryazan reg. (Negrobov, Pogonin, 1984, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Adygea (Grichanov, 2012c), Chrysotimus spinuliferus Negrobov 1978 – Primorye , Yakutia (Negrobov, 1978 b).
Chrysotus Meigen, 1824.
Chrysotus amurensis Negrobov, 1980 – Amur reg. (Negrobov, 1980 b, Maslova, Negrobov, Selivanova, 2011 b), Chrysotus andrei Negrobov, 1986 – Amur reg. (Negrobov, 1986 a), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Chrysotus angulicornis Kowarz, 1874 – Krasnodar Territory, Karachai-Cherkess Republic, Dagestan (Maslova, Negrobov, Selivanova, 2007), Chrysotus arcticus Frey, 1915 – Murmansk reg. (Frey, 1915, Grichanov, 2004), Chrysotus baicalensis Negrobov et Maslova 1995 – Irkutsk reg. (Negrobov, Maslova, 1995 b), Khanty-Mansi autonomous region (Grichanov, 2010 a), Chrysotus caerulescens Negrobov 1980 – Ural, the Lower Lena, Sayan Mountains Altai, Krasnoyarsk Territory (Negrobov, 1980 b), Altai (Negrobov, Barkalov, 2009), Khanty-Mansi autonomous region (Grichanov, 2010 a), Yamal-Nenets autonomous region, Altai, Krasnoyarsk Territory, Buryatia, Amur reg. (Maslova, Negrobov, Selivanova, 2011 b), Chrysotus cilipes Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Adygea (Grichanov, Kustov, Volfov, 2006), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Krasnodar Territory (Maslova, Negrobov, Selivanova, 2007), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Kaluga reg. (Grichanov, 2010b), Karelia, Leningrad reg., Krasnodar Territory, Altai, Krasnoyarsk Territory, Irkutsk reg., Buryatia, Baikal reg., Yakutia, Amur reg., Khabarovsk Territory, Primorye (Maslova, Negrobov, Selivanova, 2011 b), Chrysotus collini Parent 1923 – North Caucasus (Grichanov, Volfov, Kustov, 2006), Chrysotus corniger Negrobov et Maslova 1995 – Primorye (Negrobov, Maslova, 1995 b), Chrysotus cupreus Macquart, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), North Caucasus (Negrobov, 1967 d), Voronezh reg. (Negrobov, 1972), Krasnodar Territory (Maslova, Negrobov, Selivanova, 2007), Chrysotus decipiens Negrobov et Zurikov 2000 – Ural (Negrobov, Tsurikov, Maslova, 2000), Chrysotus defensus Negrobov et Maslova 2000 – Krasnodar Territory, North Caucasus (Negrobov, Tsurikov, Maslova, 2000, Maslova, Negrobov, Selivanova, 2007), Chrysotus degener Frey, 1917 – Jewish autonomous region (Grichanov, 2006 a), Chrysotus femoratus Zetterstedt 1843 – Murmansk reg. (Frey¸ Lundsrom, 1913), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c, Maslova, Negrobov, Selivanova, 2008), Yakutia (Negrobov, Chalaya, 1991), Adygea (Grichanov, Kustov, Volfov, 2006), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Krasnodar Territory (Maslova, Negrobov, Selivanova, 2007), Ryazan reg. (Negrobov, Pogonin, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Chrysotus gramineus (Fallen, 1823) (synonym Chrysotus microcerus Kowarz, 1874, Chrysotus varians Kowarz, 1874) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko A., 1891, Fedtchenko B., 1892), Murmansk reg. (Frey¸ Lundsrom, 1913, Frey, 1915, Negrobov, 1974), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 а), North Caucasus (Negrobov, 1967 c, 1968 b, Grichanov, Volfov, Kustov, 2006, 2007), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Yakutia (Negrobov, Chalaya, 1991), Krasnoyarsk Territory (Pogonin, Negrobov, 2008, Grichanov, Kustov, Volfov, 2006), Vologda reg. (Grichanov, 2006 b), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Krasnodar Territory (Maslova, Negrobov, Selivanova, 2007, Grichanov, 2012 c), Karelia, Arkhangelsk reg., Leningrad reg., Novgorod reg., Pskov reg., Yaroslavl reg., Moscow reg., Ryazan reg., Kursk reg., Lipetsk reg., Voronezh reg., Krasnodar Territory, Adygea, Karachai-Cherkess Republic, Bashkortostan, Orenburg reg., Chelyabinsk reg., Tomsk reg., Altai, Krasnoyarsk Territory, Irkutsk reg., Buryatia, Chita reg., Yakutia, Amur reg., Khabarovsk Territory, Yakutia (Maslova, Negrobov, Selivanova, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Chrysotus fuscoluteus Negrobov et Zurikov 1986 – Yakutia (Negrobov, Tsurikov, 1986 a, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Chrysotus glebi Negrobov et Maslova 1995 – Primorye (Negrobov, Maslova, 1995 b), Chrysotus laesus (Wiedemann, 1817) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Irkutsk reg. (Frey, 1915, Maslova, Negrobov, Selivanova, 2010), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 а), Krasnodar Territory (Negrobov, Gоlubcova, 1983, Maslova, Negrobov, Selivanova, 2007), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Amur reg. (Negrobov, 1986 a, 1986 b), Adygea (Grichanov, Kustov, Volfov, 2006), North Caucasus (Volfov, Kustov, 2006), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Karelia, Pskov reg., Yaroslavl reg., Moscow reg., Kursk reg., Lipetsk reg., Voronezh reg., Krasnodar Territory, Tatarstan, Bashkiria, Orenburg reg., Omsk reg., Tomsk reg., Altai, Krasnoyarsk Territory, Irkutsk reg., Buryatia, Yakutia, Amur reg. (Maslova, Negrobov, Selivanova, 2011 b), Chrysotus ljutengensis Negrobov et Zurikov 1986 – Yakutia (Negrobov, Tsurikov, 1986 a, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Chrysotus logvinovskii Negrobov et Zurikov 2000 – Yakutia (Negrobov, Tsurikov, 2000, Negrobov, Tsurikov, Maslova, 2000), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Chrysotus neglectus (Wiedemann, 1817) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko A., 1891, Fedtchenko B., 1892), Volga River basin (Eversmann, 1834), Murmansk reg. (Frey, 1915), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), Kamchatka (Parent, 1930), Voronezh reg. (Negrobov, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Ryazan reg. (Negrobov, Pogonin, 1984), Amur reg. (Negrobov, 1986 b), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Adygea (Grichanov, Kustov, Volfov, 2006), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Krasnodar Territory (Maslova, Negrobov, Selivanova, 2007), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Dagestan (Grichanov, 2012 c), Chrysotus nudisetus Negrobov et Maslova 1995 – Yakutia (Negrobov, Maslova, 1995 b, Grichanov, Bagachanova, 2006), Chrysotus obscuripes Zetterstedt 1838 (synonym – Chrysotus kowarzi Lundbeck, 1912)
– Leningrad reg. (Stackelberg, 1962), Yakutia (Grichanov, Bagachanova, 2006), Chrysotus orientalis Negrobov et Zurikov 2000 – Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Chrysotus parilis Parent 1926 – Baikal, Primorye (Maslova, Negrobov, Selivanova, 2011 b), Chrysotus peculiariter Negrobov et Maslova 2000 – North Caucasus (Negrobov, Tsurikov, Maslova, 2000, Grichanov, Volfov, Kustov, 2007), Chrysotus pennatus Lichtwardt 1902 – Krasnodar Territory (Negrobov, 1965b, Maslova, Negrobov, Selivanova, 2007, Grichanov, 2012 c), Voronezh reg. (Negrobov, 1967 а), Adygea (Grichanov, Kustov, Volfov, 2006), Adygea, Krasnodar Territory, Adygea, North Caucasus (Maslova, Negrobov, Selivanova, 2011b), Chrysotus pilitibia Negrobov et Maslova 1995 – Ural (Negrobov, Maslova, 1995 b), Chrysotus pseudocilipes Hollis 1964 – Amur reg., Primorye (Maslova, Negrobov, Selivanova, 2011 b), Chrysotus pulchellus Kowarz 1874 – Murmansk reg. (Frey, 1915), Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1967 a), North Caucasus (Negrobov, 1967 c), Ryazan reg. (Negrobov, Pogonin, 1984), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Yakutia (Negrobov, Chalaya, 1991), Adygea (Grichanov, Kustov, Volfov, 2006), Krasnodar Territory (Maslova, Negrobov, Selivanova, 2007), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Chrysotus sibiricus Negrobov et Maslova 1995 – Far East (Negrobov, Maslova, 1995 b), Chrysotus smithi Negrobov 1980 – Baikal (Negrobov, 1980 b), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Chrysotus suavis Loew 1857 – Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Krasnodar Territory (Negrobov, Duhanina, 1984, Grichanov, 2012 c), Ryazan reg. (Negrobov, Pogonin, 1984), Yakutia (Negrobov, Chalaya, 1991), Adygea (Grichanov, Kustov, Volfov, 2006, Maslova, Negrobov, Selivanova, 2007), Altai (Grichanov, 2007b, Negrobov, Barkalov, 2009), Ryazan reg. (Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Leningrad reg., Kursk reg., Lipetsk reg., Voronezh reg., Rostov reg., Adygea, Altai, Krasnoyarsk Territory, Irkutsk reg., Buryatia, Yakutia, Amur reg., Magadan reg., Khabarovsk Territory, Kamchatka, Yakutia обл. (Maslova, Negrobov, Selivanova, 2011 b), Astrakhan reg. (Grichanov, 2011), Chrysotus viridifemoratus Roser, 1840 (synonym – Chrysotus monochaetus Kowarz 1874) – Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Ryazan reg. (Negrobov, Pogonin, 2008), Khanty-Mansi autonomous region (Grichanov, 2010 a), Chrysotus vladimiri Megrobov et Maslova 1995 – Amur reg. (Negrobov, Maslova, 1995 b), Chrysotus zlobiniani Negrobov et Maslova 1995 – Yakutia (Negrobov, Maslova, 1995 b).
Diaphorus Meigen, 1824.
Diaphorus anatoli Negrobov 1986 – Amur reg. (Negrobov, 1976 a, 1986 b), Diaphorus andreji Negrobov 1976 – Amur (Negrobov, 1976 a), Diaphorus deliquescens Loew 1871 – Moscow reg. (Fedtchenko A., 1892), Central Russia (Stackelberg, 1928 b), Leningrad reg. (Stackelberg, 1962), North Caucasus (Grichanov, Volfov, Kustov, 2007), Diaphorus disjunctus Loew 1857 – Leningrad reg. (Stackelberg, 1962), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2006, Grichanov, Volfov, Kustov, 2007), Adygea (Grichanov, Volfov, Kustov, 2009), Diaphorus exunguiculatus Parent 1925 – Leningrad reg. (Stackelberg, 1925, 1928, 1962), Siberia (Negrobov, Maslova, Selivanova, 2005 a), Diaphorus hoffmannseggi Meigen, 1830 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Ural (Frеу, 1915), Leningrad reg. (Stackelberg, 1925, 1928, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c), Diaphorus oculatus (Fallen, 1823) – Ryazan reg. (Negrobov, Pogonin, 2008), Diaphorus nigricans Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), North Caucasus (Negrobov, 1967 c), Krasnodar Territory (Negrobov, Duhanina, 1984), Amur reg. (Negrobov, 1986 b), Murmansk reg. (Grichanov, 2004), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Diaphorus oculatus (Fallén, 1823) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1972), Ryazan reg. (Negrobov, Pogonin, 1984), North Caucasus (Grichanov, Volfov, Kustov, 2007), Diaphorus parenti Stackelberg 1928 – Primorye (Stackelberg, 1928 b, Negrobov, Maslova, 2005), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Diaphorus pilitibius Negrobov et Maslova 1995 – Krasnodar Territory (Negrobov, Maslova, 1995), North Caucasus (Grichanov, Volfov, Kustov, 2006, Grichanov, Volfov, Kustov, 2007), Diaphorus sokolovi – Stackelberg, 1928 – Primorye, Chita reg. (Stackelberg, 1928 b), Diaphorus ussuriensis Stackelberg 1928 – Primorye , Chita reg. (Stackelberg, 1928 b), Primorye (Negrobov, Maslova, 2005), Diaphorus vitripennis Loew, 1859 – Orenburg reg. (Beker, 1918, Wnukowsky, 1936), Primorye (Stackelberg, 1928 b), Russia (Parent, 1938), Diaphorus winthemi Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), – Orenburg reg. (Beker, 1918, Wnukowsky, 1936), Primorye (Stackelberg, 1928 b), Russia (Parent, 1938), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Diaphorus zlobini Negrobov et Duchanina 1987 – Sakhalin (Negrobov, Duhanina, 1987).
Diostracus Loew, 1861.
Diostracus maculatus Negrobov 1980 – Primorye (Negrobov, 1980 a), Diostracus naegelei Negrobov 1978 – Yakutia (Negrobov, 1978 b), Diostracus subalpinus Negrobov 1973 – Buryatia (Negrobov, 1973 d), Diostracus vitae Negrobov 1980 – Primorye (Negrobov, 1980 a), Diostracus zlobini Negrobov 1980 – Primorye (Negrobov, 1980 a).
Dolichophorus Lichtwardt, 1902
Dolichophorus kerteszi Lichtwardt, 1902 – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938).
Dolichopus Latreille, 1796.
Dolichopus acuticornis Wiedemann 1817 – Moscow reg. (Eversmann, 1834), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Voronezh reg. (Negrobov, 1967 a), Ryazan reg. (Negrobov, Pogonin, 1984), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Dolichopus agilis Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko A., 1891, Fedtchenko B., 1892), Siberia (Stackelberg, 1930 a), Far East (Stackelberg, 1930 b), Russia (Parent, 1938), Sayan Mountains (Negrobov, 1973 a, Negrobov, Selivanova, Maslova, 2010), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Buryatia, Irkutsk reg. (Negrobov, Radionova, 2004), Jewish AR, Khabarovsk Territory (Grichanov, 2006 a), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Ryazan reg. (Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Dolichopus albicinctus Smirnov 1948 – Primorye (Smirnov, 1948 а, Negrobov, Sviridova, 1983), Dolichopus altayensis Yang, 1998 – Altai (Grichanov, 2007 , Negrobov, Barkalov, 2009), Dolichopus amginensis Stackelberg 1928 – Yakutia (Stackelberg, 1928 a, 1929, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Siberia (Stackelberg, 1930 a), Dolichopus amurensis Stackelberg 1930 – Amur reg. (Stackelberg, 1930 a), Primorye (Negrobov, Sviridova, 1983), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Khabarovsk Territory (Grichanov, 2006 a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Altai (Negrobov, Barkalov, 2009), Khabarovsk Territory, Amur reg. (Kornev, Negrobov, Maslova, 2011), Dolichopus angustipennis Kertész, 1901 – Karelia (Frey, 1915), Siberia (Stackelberg, 1930), Leningrad reg. (Stackelberg, 1962), Buryatia (Negrobov, Rodionova, 2004 а), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Irkutsk reg., Свердловская обл. (Kornev, Negrobov, Maslova, 2011), Dolichopus annulipes Zetterstedt, 1938 – Moscow reg. (Fedtchenko A., 1868), Murmansk reg. (Frey, 1915), Ural (Becker, 1915, Negrobov, Rodionova, 2004 а), Yakutia (Stackelberg, 1930 a, Grichanov, Bagachanova, 2006), Leningrad reg. (Stackelberg, 1962), Murmansk reg. (Negrobov, 1968 b), Murmansk reg. (Negrobov, 1974), Khanty-Mansi autonomous region (Grichanov, 2010 a), Dolichopus annulitarsis Ringdahl 1920 – Ural (Stackelberg, 1930 a), Magadan reg. (Negrobov, Chalaya, 1991), Yakutia (Grichanov, Bagachanova, 2006), Dolichopus apicalis Zetterstedt 1849 – Novgorod reg. (Stackelberg, 1919), Far East (Stackelberg, 1930 a), Voronezh reg. (Negrobov, 1967 a), Kamchatka (Negrobov, Rodionova, 2004 а), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Vologda reg. (Grichanov, 2006 b), Kursk reg. (Grichanov, 2007 a), Dolichopus arbustorum Stannius 1831 – North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006), Voronezh reg. (Negrobov, 1972), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Ryazan reg. (Negrobov, Pogonin, 2008), Dolichopus argyrotarsis Wahlberg 1850 – Leningrad reg. (Stackelberg, 1925, 1962), Siberia (Stackelberg, 1930 a), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), North Caucasus (Grichanov, Volfov, Kustov, 2007), Dolichopus armillatus Wahlberg, 1850 – Murmansk reg. (Frey, 1915), Ural (Stackelberg, 1930 a), Dolichopus asiaticus Negrobov 1973 – Buryatia (Negrobov, 1973 a), Dolichopus atripes Meigen, 1824 Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1922, 1925, 1962), Dolichopus atritibialis Zetterstedt 1859 – Arkhangelsk reg. (Negrobov, 1977 b), Dolichopus austriacus Parent 1927 – Volograd reg. (Stackelberg, 1930 a), European part of Russia (Negrobov, 1977b), Astrakhan reg. (Grichanov, 2011), Dolichopus basalis Loew 1859 – Kamchatka (Stackelberg, 1930 a), Magadan reg. (Negrobov, Chalaya, 1991), Dolichopus bianchii Stackelberg 1929 – North Ural (Stackelberg, 1929), Primorye (Stackelberg, 1930 a, Negrobov, Sviridova, 1983), Yakutia (Negrobov, 1973 c, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Dolichopus bigeniculatus Parent 1926 – Primorye (Stackelberg, 1930 b, Negrobov, Sviridova, 1983, Stackelberg, 1930, Udovenko, 1970), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus bilamellatus Parent, 1929 – Russia (Parent, 1929), Primorye (Stackelberg, 1930 a, Stackelberg, 1930 b), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus bisetulatus Negrobov 1977 – Primorye (Negrobov, 1977), Dolichopus bonsdorfii Frey 1915 – Leningrad reg. (Stackelberg, 1962, Negrobov, 1977 b), Khabarovsk Territory (Negrobov, 1977 b), Altai (Negrobov, Barkalov, 2009), Dolichopus breviclypeus Negrobov, 1976 –Buryatia (Negrobov, 1976 c), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Dolichopus brevipennis Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtscenko A., 1891, Fedtchenko B., 1892), Murmansk reg. (Frey¸ Lundsrom, 1913, Negrobov, 1974), Leningrad reg. (Stackelberg, 1921, 1925), Yakutia (Stackelberg, 1928 a, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Far East (Stackelberg, 1930 a), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Krasnodar Territory (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006), Altai (Grichanov, 2007b, Negrobov, Barkalov, 2009), Kabardino-Balkaria, Adygea (Grichanov, Volfov, Kustov, 2007), Khanty-Mansi autonomous region (Grichanov, 2010 a), Dolichopus calceatus Parent, 1927 – Transbaikale (Parent, 1927), Chita reg. (Stackelberg, 1930 a), Primorye (Stackelberg, 1930 b), Khabarovsk reg. (Negrobov, Sviridova, 1983), Kamchatka (Negrobov, Rodionova, 2004 а), Yakutia (Grichanov, Bagachanova, 2006), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Dolichopus caligatus Wahlberg, 1850 – Khabarovsk Territory (Grichanov, 2006 a), Dolichopus calinotus Loew 1871 – North Caucasus (Grichanov, Volfov, Kustov, 2006), Dolichopus campestris Meigen 1824 – Siberia (Frey 1915), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), Primorye (Stackelberg, 1930 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Kamchatka (Negrobov, Chalaya, 1991), Vologda reg. (Grichanov, 2006 b), Altai (Grichanov, 2007 b), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Dolichopus cilifemoratus Macquart 1827 – Moscow reg. (Fedtchenko A., 1891, Fedtchenko B., 1892), Siberia (Frey, 1915, Stackelberg, 1930), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1922, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1967 a, 1972), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006), Tomsk reg. (Negrobov, Rodionova, 2004 а), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Ryazan reg. (Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Dolichopus cinctipes Wahlberg 1850 – Siberia (Stackelberg, 1930 a), Baikal (Negrobov, 1977 b), Arkhangelsk reg. (Negrobov, Rodionova, 2004 а), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Komi, Chukchi autonomous Region (Grichanov, 2012 b), Dolichopus ciscaucasicus Stackelberg 1927 – North Caucasus (Stackelberg, 1927, Stackelberg, 1930, Negrobov, 1965 b, 1965, 1967 c), Dolichopus claviger Stannius 1831 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1982), Siberia (Frey, 1915), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Siberia (Stackelberg, 1930 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Magadan reg. (Negrobov, Chalaya, 1991), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Vologda reg. (Grichanov, 2006b), Yakutia (Grichanov, Bagachanova, 2006), Altai (Grichanov, 2007b), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Altai (Negrobov, Barkalov, 2009), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Dolichopus clavipes clavipes Yaliday, 1832 – Murmansk reg. (Negrobov, 1974), Murmansk reg., Baikal region (Negrobov, Булли, 1986), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Urals, Yakutia, Tomsk reg. (Negrobov, Rodionova, 2004 а), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Dolichopus clavipes fusiformis Becker, 1917 (Turkestan) – Yakutia (Stackelberg, 1930 a, Negrobov, Chalaya, 1991), Dolichopus costalis Frey, 1915 – Russia (Frey, 1915), Karelia (Stackelberg, 1929), Yakutia (Stackelberg, 1930 a, Negrobov, 1973 a), Dolichopus cruralis Wahlberg, 1850 – Leningrad reg. (Stackelberg, 1925, 1962), Siberia (Stackelberg, 1930 a), Russia (Parent, 1938), Murmansk reg. (Grichanov, 2004), Dolichopus czekanovskii Stackelberg 1928 – Yakutia (Stackelberg, 1928 a, 1930, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Dolichopus davshinicus Negrobov 1973 – Buryatia (Negrobov, 1973 b), Yakutia (Negrobov, Chalaya, 1991), Dolichopus diadema Haliday, 1832 – North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Dolichopus discimanus Wahlberg, 1851 – Siberia (Stackelberg, 1930 a), Primorye (Negrobov, Sviridova, 1983), Dolichopus divisus Becker 1917 – Far East (Negrobov, Sviridova, 1983), Altai (Negrobov, Barkalov, 2009), Dolichopus disharmonicus Smirnov, 1948 – Primorye (Smirnov, 1948 а), Dolichopus divisus Becker, 1917 – Ural (Becker, 1917), Irkutsk reg. (Stackelberg, 1930 a), Dolichopus emelijanovi Stackalberg, 1930 – Primorye (Stackelberg, 1930 a, Stackelberg, 1930 b), Dolichopus eous Stackelberg 1929 – Buryatia (Stackelberg, 1929), Yakutia (Stackelberg, 1930 a, Grichanov, Bagachanova, 2006), Buryatia (Negrobov, Rodionova, 2004 а), Dolichopus eurypterus Gerstacker 1864 – Primorye (Stackelberg, 1930 a, Stackelberg, 1930 b), Khabarovsk Territory (Grichanov, 2006a), Dolichopus excisus Loew 1859 – Siberia (Stackelberg, 1930 a), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1972), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Adygea (Grichanov, Kustov, Volfov, 2006), Krasnodar Territory (Grichanov, 2012 c), Dolichopus flavipes Stannius 1831 – Siberia (Frey, 1915,), Leningrad reg. (Stackelberg, 1925, 1962), Yakutia (Stackelberg, 1930 a, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Primorye (Stackelberg, 1930 a), Russia (Parent, 1938), Far East (Negrobov, Sviridova, 1983), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Dolichopus fraterculus fraterculus Zetterstedt 1843 – Murmansk reg. (Frey, 1915, Negrobov, 1974), Karelia (Stackelberg, 1930 a), Yakutia (Grichanov, Bagachanova, 2006), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Dolichopus fraterculus nigrifemur Stackelberg, 1930 –Ural (Stackelberg, 1930), Dolichopus fridolini Stackelberg, 1928 – Ural (Stackelberg, 1928 a, 1930), Dolichopus fursovi Negrobov et Barkalov, 2010 – Altai (Negrobov, Barkalov, 2010), Dolichopus galeatus Loew, 1871 – "Sibirien" (Loew, 1871), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Kamchatka (Parent, 1930), Primorye (Stackelberg, 1930 a), Khabarovsk Territory (Negrobov, Sviridova, 1983), Dolichopus gorodkovi Negrobov 1973 – Sayan Mountains, Irkut reg. (Negrobov, 1973 b), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Dolichopus grandicornis Wahlberg, 1850 – Ural (Becker, 1917), Dolichopus griseifacies Becker, 1917 – Irkutsk (Becker, 1917), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus griseipennis Stannius, 1831 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1982), North Caucasus (Stackelberg, 1926 а, Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Siberia (Stackelberg, 1930 a), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), Dolichopus grunini Smirnov, 1948 – Primorye (Smirnov, 1948 а), Magadan reg. (Negrobov, Chalaya, 1991), Dolichopus gubernator Mik, 1978 – Leningrad reg. (Stackelberg, 1962), Khabarovsk Territory (Negrobov, 1977 b), Dolichopus hilaris Loew, 1862 – Primorye (Stackelberg, 1930 a, Stackelberg, 1930 b), Russia (Parent, 1938), Khabarovsk Territory (Negrobov, 1977 b), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Dolichopus humilis Van Duzee, 1921 (synonym Dolichopus lanzovi Negrobov, Grichanov et Barkalov 2009) – Kamchatka (Negrobov, Rodionova, 2004а), Altai, Yakutia, Irkutsk reg. (Negrobov, Grichanov, Barkalov, 2009), Taimyr (Negrobov, Grichanov, Barkalov, 2009), Yakutia (Grichanov, Bagachanova, 2006), Dolichopus impotens Smirnov, 1948 – Primorye (Smirnov, 1948b), Dolichopus intonsus Smirnov, 1948 – Primorye (Smirnov, 1948 b), Dolichopus ivanovi Stackelberg 1929 – Yakutia (Stackelberg, 1929 a, 1930, Grichanov, Bagachanova, 2006), Kamchatka (Negrobov, Chalaya, 1991), Dolichopus jacutensis Stackelberg 1929 – Yakutia (Stackelberg, 1929 a, 1930, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006, Kornev, Negrobov, Maslova, 2011), Magadan reg., Khabarovsk reg., Primorye (Negrobov, Sviridova, 1983, Maslova, Negrobov, Kornev, 2011), Dolichopus jakutus Selivanova et Negrobov, 2011 – Yakutia (Selivanova, Negrobov, 2011), Dolichopus kjari Stackelberg 1929 – Yakutia (Stackelberg, 1929 a, 1930, Negrobov, 1973 a, Grichanov, Bagachanova, 2006), Irkutsk reg. (Stackelberg, 1929, 1930, Negrobov, 1973 c), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Dolichopus kurayensis Negrobov, Barkalov et Selivanova 2011 – Altai (Negrobov, Barkalov, Selivanova, 2011), Dolichopus kuznetsovi, Maslova, Negrobov, Selivanova, 2012 – Primorye (Maslova, Negrobov, Selivanova, 2012), Dolichopus lancearius Hedström 1966 – Baikal (Negrobov, 1976 c), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Dolichopus latilimbatus Macquart 1827 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Yakutia (Stackelberg, 1930 a), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, Negrobov, 1965 c, 1966 а, 1967 a, 1972), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Krasnodar Territory (Negrobov, Duhanina, 1984, Grichanov, Kustov, Volfov, 2006), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Kursk reg. (Grichanov, 2007 a), Dolichopus latipennis Fallen, 1823 – Transbaikalia, Yakutia (Stackelberg, 1933), Russia (Paren, 1938), Murmansk reg. (Negrobov, 1974), Dolichopus lepidus lepidus Staeger, 1842 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915, Negrobov, 1974), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1967a), North Caucasus (Negrobov, 1967 c, 1970), Magadan reg. (Negrobov, Chalaya, 1991), Khanty-Mansi autonomous region (Grichanov, 2010 a) , Adygea (Negrobov, Radionova, 2004), Vologda reg. (Grichanov, 2006 b), Khabarovsk Territory (Grichanov, 2006a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Dolichopus lepidus microstigma Stackelberg 1930 – Primorye (Stackelberg, 1930 b, Stackelberg, 1930 a, Negrobov, Sviridova, 1983), Ryazan reg. (Negrobov, Pogonin, 2008), Dolichopus leucopus Smirnov, 1948 – Primorye (Smirnov, 1948 b), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus linearis Meigen, 1824
– Siberia (Frey, 1915, Negrobov, 1973 a), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1922, 1925, 1962), Primorye (Stackelberg, 1930 a, Stackelberg, 1930 b, Negrobov, Sviridova, 1983), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1967 a), North Caucasus (Negrobov, 1967 c), Ryazan reg. (Negrobov, Pogonin, 1984), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Khanty-Mansi autonomous region, Adygea (Negrobov, Radionova, 2004), Vologda reg. (Grichanov, 2006 b), Khabarovsk Territory (Grichanov, 2006 a), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Ryazan reg. (Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Khanty-Mansi autonomous region (Grichanov, 2010 a), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Dolichopus lineaticornis Zetterstedt, 1843 – Krasnodar Territory (Negrobov, Duhanina, 1984), Ryazan reg. (Negrobov, Pogonin, 1984), Dolichopus litorellus Zetterstedt, 1852 – Moskov reg., Yakutia (Stackelberg, 1933), Yakutia (Negrobov, 1973 a, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Omsk reg. (Negrobov, Rodionova, 2004 а), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Dolichopus lonchophorus Loew, 1873 – “Kultuk” [= near Irkutsk] (Loew, 1873). Siberia (Stackelberg, 1928 a, Negrobov, 1973 a), Ural (Stackelberg, 1933, Negrobov, 1973 c), Khabarovsk Territory (Grichanov, 2006 a), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Dolichopus longicornis Stannius, 1831 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko A., 1891, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915, Negrobov, 1968 b, Frey¸ Lundsrom, 1913), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1922, 1925, 1962), Far East (Stackelberg, 1930 b), Moscow reg., Perm reg., Irkutsk reg., Kamchatka (Stackelberg, 1933), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c), Primorye (Negrobov, Sviridova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Yakutia (Negrobov, 1973 a, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Vologda reg. (Grichanov, 2006 b), Altai (Grichanov, 2007 b), Kursk reg. (Grichanov, 2007 a), Ryazan reg. (Negrobov, Pogonin, 2008), Kaluga reg. (Grichanov, 2010 b), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Dolichopus longicostalis Negrobov et Barkalov, 1978 – Yakutia (Negrobov, Barkalov, 1978), Dolichopus longisetus Negrobov, 1977 – Primorye (Negrobov, 1977 c), Khabarovsk Territory (Grichanov, 2006 a), Altai (Grichanov, 2007 b), Dolichopus longitarsis Stannius, 1831 – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Siberia (Stackelberg, 1928 a), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1967 а), Yakutia (Negrobov, Barkalov, 1978), Vologda reg. (Grichanov, 2006 b), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Altai (Negrobov, Barkalov, 2009), Dolichopus luteifacies Parent, 1927 – Irkutsk reg. (Parent, 1927), Primorye (Stackelberg,1933, Udovenko, 1970), Dolichopus maculicornis Verrall, 1875 – Northern Europe Yakutia (Grichanov, Bagachanova, 2006), Dolichopus maculipennis Zetterstedt, 1843 – Murmansk reg. (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Kamchatka (Parent, 1930), Northern Europe (Stackelberg, 1933), Magadan reg. (Negrobov, Chalaya, 1991), Yakutia (Grichanov, Bagachanova, 2006), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Dolichopus makarovi Smirnov, 1948 – Primorye (Smirnov, 1948 b, Negrobov, Sviridova, 1983), Kamchatka (Negrobov, Rodionova, 2004 а), Dolichopus mannerheimi Zetterstedt, 1838 – Murmansk reg. (Frey, 1915), Siberia (Negrobov, 1973 a), Khanty-Mansi autonomous region (Negrobov, 1973 c, Grichanov, 2010 a), Altai, Irkutsk reg., Transbaikalia, Khabarovsk Territory, Primorye, Magadan reg., Kamchatka (Negrobov, 1977b), Magadan reg. (Negrobov, Chalaya, 1991), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Dolichopus martynovi Stackelberg, 1930 – Primorye (Stackelberg, 1930 b, 1933, Negrobov, 1973 c), Dolichopus mediovenus Negrobov, 1977 – Primorye (Negrobov, 1977), Murmansk reg. (Grichanov, 2004), Dolichopus meigeni Loew 1857 – Voronezh reg. (Negrobov, 1977 g, Negrobov, Radionova, 2004), Dolichopus melanopus Meigen 1824 – Ural (Eversmann, 1834), Dolichopus migrans Zetterstedt, 1843 – Karelia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Far East (Stackelberg, 1930 a), Siberia (Stackelberg, 1933), Voronezh reg. (Negrobov, 1963, 1965 c, 1966 а, 1967 a), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Magadan reg. (Negrobov, Chalaya, 1991), Dolichopus monochaetus Smirnov, 1948 – Primorye (Smirnov, 1948b), Dolichopus nataliae Stackelberg, 1930 – Primorye (Stackelberg, 1933), Magadan reg. (Negrobov, Chalaya, 1991), Yakutia (Grichanov, Bagachanova, 2006), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus nebulosus Smirnov, 1948 – Primorye (Smirnov, 1948 b), Dolichopus negrobovi Gosseries, 1989 (synonym – Dolichopus pallipes Negrobov, 1973) – Buryatia (Negrobov, 1973 a), Magadan reg. (Negrobov, Chalaya, 1991), Yakutia (Grichanov, Bagachanova, 2006), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus nigrilamellatus Becker, 1917 – Ural (Becker, 1917), Dolichopus nigricornis Meigen, 1824 (synonym – Dolichopus discifer Stannius, 1831) – Moscow reg. (Fedtchenko B., 1892), Murmansk reg. (Frey, 1915, Negrobov, 1968 b, 1974), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Far East (Negrobov, Sviridova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Magadan reg. (Negrobov, Chalaya, 1991), Yamal-Nenets autonomous region (Negrobov, Rodionova, 2004а), Vologda reg. (Grichanov, 2006 b), Yakutia (Grichanov, Bagachanova, 2006), Khabarovsk Territory (Grichanov, 2006 a), Altai (Grichanov, 2007 b), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Khanty-Mansi autonomous region (Grichanov, 2010 a), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Dolichopus nirgipes Fallen, 1823 – Voronezh reg. (Negrobov, 1972, 1977 b, Negrobov, 1977 g), Dolichopus nitidus Fallén, 1823 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), Far East (Stackelberg, 1930 b), Siberia (Stackelberg, 1933), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1965 с, 1967 a), Primorye (Udovenko, 1970), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Khanty-Mansi AR (Grichanov, 2010a), North Caucasus (Grichanov, Kustov, Volfov, 2006), Khabarovsk Territory (Grichanov, 2006a), Altai (Grichanov, 2007b), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Dolichopus notatus Staeger, 1842 – Moscow reg. (Fedtchenko B., 1892), Karelia (Frey, 1915), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Siberia (Stackelberg, 1933), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1967 a), North Caucasus (Negrobov, 1970 b), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Magadan reg. (Negrobov, Chalaya, 1991), Yakutia (Grichanov, Bagachanova, 2006), Altai (Grichanov, 2007 b), Dolichopus nubilus Meigen, 1824 – Karelia (Frey, 1915), Siberia (Stackelberg, 1933), Russia (Parent, 1938), Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1972), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Kursk reg. (Grichanov, 2007 a), Altai (Negrobov, Barkalov, 2009), Dolichopus ornatitarsis Negrobov Barkalov, 2008 – Altai (Negrobov, Barkalov, 2009), Dolichopus pamiricus Negrobov, 1976 – Pamir (Negrobov, 1976 c), Dolichopus pectinitarsis Stenhammer, 1851 – North Ural (Stackelberg, 1933), Yakutia (Grichanov, Bagachanova, 2006), Dolichopus pennatus Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko A., 1891, Fedtchenko B., 1892), Murmansk reg. (Frey¸ Lundsrom, 1913, Novgorod reg. (Stackelberg, 1919), Pskov reg. (Stackelberg,1933), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Magadan reg. (Negrobov, Chalaya, 1991), Vologda reg. (Grichanov, 2006 b), Yakutia (Grichanov, Bagachanova, 2006), Altai (Grichanov, 2007 b), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Dolichopus picipes Meigen, 1824 – Karelia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Mosow reg. (Stackelberg, 1933), Voronezh reg. (Negrobov, 1967 a, 1972), North Caucasus (Negrobov, 1967 c, 1968 b), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Altai (Grichanov, 2007 b), Ryazan reg. (Negrobov, Pogonin, 2008), Dolichopus planitarsis Fallén, 1823 – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962, 1933), Voronezh reg. (Negrobov, 1967 a), Kamchatka (Negrobov, Rodionova, 2004 а), Yakutia (Grichanov, Bagachanova, 2006), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Altai (Barkalov, Negrobov, Grichanov, 2009), Dolichopus platychaetus Negrobov et Barkalov, 1977 – Magadan reg. (Negrobov, Barkalov, 1977, Negrobov, Chalaya, 1991), Dolichopus plumipes Scopoli, 1763 (synonym Dolichopus parvicaudatus Zetterstedt, 1843), – Moscow reg. (Fedtchenko A., 1868, Fedtchenko A., 1891, Fedtchenko B., 1892), Murmansk reg. (Frey¸ Lundsrom, 1913, Ural (Becker, 1915), Murmansk reg. (Frey, 1915, Negrobov, 1968 b, Negrobov, 1974), Russia (Frey, 1918), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Far East (Stackelberg, 1930 a, Negrobov, Sviridova, 1983), Kamchatka (Parent, 1930 b), Siberia (Stackelberg, 1933), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Primorye (Udovenko, 1970), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Kamchatka, Yakutia (Negrobov, Rodionova, 2004а), Khabarovsk Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2006a), Vologda reg. (Grichanov, 2006b), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Kursk reg. (Grichanov, 2007 a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Khanty-Mansi autonomous region (Grichanov, 2010 a), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Krasnodar Territory (Grichanov, 2012 c), Dolichopus plumitarsis Fallén, 1823 – Volga River basin (Eversmann, 1834), Moscow reg. (Fedtchenko A., 1868), Far East (Stackelberg, 1930 b), Perm reg., Irkutsk reg, Primorye (Stackelberg, 1933), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c), Primorye (Udovenko, 1970, Negrobov, Sviridova, 1983), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Tomsk reg., Altai (Negrobov, Rodionova, 2004 а), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Altai (Negrobov, Barkalov, 2009), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Dolichopus polychaetus Negrobov, 1973 (Mongolia) – Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Dolichopus popularis Wiedemann, 1817 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892, Stackelberg, 1933), Siberia (Frey, 1915), Leningrad reg. (Porchinskij, 1874, Stackelberg, 1925, 1962), Russia (Parent, 1928), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Altai (Grichanov, 2007 b), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Dolichopus portentosus Negrobov, 1973 – Irkut reg., Buryatia (Negrobov, 1973 b), Yakutia (Negrobov, Chalaya, 1991), Dolichopus pospelovi Smirnov, 1948 – Primorye (Smirnov, 1948 b), Yakutia (Grichanov, Bagachanova, 2006), Dolichopus postocularis Negrobov, 1977 – Primorye, Kamchatka, Khabarovsk Territory (Negrobov, 1977 с), Dolichopus pullus Smirnov, 1948 – Primorye (Smirnov, 1948b), Dolichopus punctum Meigen, 1824 – Moscow reg. (Федчеко А., 1868), Leningrad reg. (Stackelberg, 1962), Yakutia (Grichanov, Bagachanova, 2006), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus remipes Wahlberg, 1839 – Leningrad reg. (Stackelberg, 1922, 1925, 1962), Kamchatka, Baikal, Buryatia (Negrobov, 1977b), Voronezh reg. (Negrobov, Korneev, 2010), Khanty-Mansi autonomous region (Negrobov, Korneev, 2010, Grichanov, 2010 a), Dolichopus rezvorum Stackelberg, 1930 – Primorye (Stackelberg, 1930 b, 1933, Udovenko, 1970), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus ringdahli Stackelberg, 1929 – Siberia (Stackelberg, 1929), Yakutia (Stackelberg, 1930 b, 1933, Grichanov, Bagachanova, 2006), Far East (Negrobov, Sviridova, 1983), Yakutia (Negrobov, Chalaya, 1991), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Krasnodar Territory, Primorye (Maslova, Negrobov, Korneev, 2012), Dolichopus robustus Stackelberg, 1928 – Primorye (Stackelberg, 1928 a, Negrobov, Sviridova, 1983), Kamchatka (Stackelberg, 1930 b, Negrobov, Rodionova, 2004 а), Khabarovsk Territory (Grichanov, 2006a), Altai (Grichanov, 2007 b), Moskow reg., Yakutia, Amur reg (Stackelberg, 1933), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Dolichopus romanovi Smirnov et Negrobov, 1973 – Baikal (Negrobov, 1973 a), Dolichopus rotundipennis Loew, 1848 – Siberia (Loew, 1848), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Baikal (Negrobov, Grichanov, Barkalov, 2009), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Dolichopus rufitinctus Becker, 1917 – Sverdlovsk reg. (Becker, 1917), Dolichopus rupestris Haliday, 1833 – Siberia (Stackelberg, 1928 a, 1933), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Voronezh reg. (Negrobov, 1972), Murmansk reg. (Negrobov, 1974), Primorye (Negrobov, Sviridova, 1983), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Ural (Negrobov, Rodionova, 2004а), Altai (Grichanov, 2007 b), Dolichopus ruthei Loew, 1847 – Moscow reg. (Fedtchenko A., 1868), Dolichopus sabinus Haliday, 1838 – North Caucasus (Grichanov, Volfov, Kustov, 2007), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Dolichopus sagittarius Loew, 1848 – Siberia (Loew, 1848, Stackelberg, 1933), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Altai (Negrobov, Barkalov, 2009), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Dolichopus salictorum (Loew, 1871) – North Caucasus (Grichanov, Volfov, Kustov, 2007), Voronezh reg. (Negrobov, Korneev, 2010), Yakutia (Maslova, Negrobov, Selivanova, 2011 a), Dolichopus saxicola Smirnov, 1948 – Primorye (Smirnov, 1948 b), Far East (Negrobov, Sviridova, 1983), Dolichopus setiger Negrobov, 1973 – Buryatia (Negrobov, 1973 b), Kamchatka (Negrobov, Rodionova, 2004а), Dolichopus selivanovae Negrobov et Barkalov, 2010 – Altai (Negrobov, Barkalov, 2010), Dolichopus setimamis Smirnov, 1948 – Primorye (Smirnov, 1948 b, Negrobov, Sviridova, 1983), Khabarovsk Territory, Chita reg., Sachalin, Kuril isl. (Selivanova, Negrobov, Maslova, 2010), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus shantaricus Stackelberg, 1933 – Khabarovsk Territory (Stackelberg, 1933), Dolichopus sharovi Smirnov, 1948 – Primorye (Smirnov, 1948 b), Dolichopus sibiricus Stackelberg, 1929 – Yakutia (Stackelberg, 1929, 1933, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Dolichopus signatus Meigen, 1824 – Leningrad reg. (Stackelberg, 1922, 1925, 1962), Voronezh reg. (Negrobov, 1972), Murmansk reg. (Negrobov, 1974), Kamchatka (Negrobov, Rodionova, 2004а), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Dolichopus signifer Haliday, 1838 –Russia (Parent, 1938), North Caucasus (Negrobov, 1965, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Voronezh reg. (Negrobov, 1965 b, 1966 а), Adygea (Grichanov, Volfov, Kustov, 2009), Dolichopus simius Parent, 1927
– Irkutsk reg. (Parent, 1927, Maslova, Negrobov, Selivanova, 2010), Yakutia (Parent, 1930), Siberia (Stackelberg, 1933, Negrobov, 1973 a), Primorye (Udovenko, 1970, Negrobov, 1973 c), Buryatia (Negrobov, Sviridova, 1983, Negrobov, Rodionova, 2004 а), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Khabarovsk Territory (Grichanov, 2006 a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Altai (Negrobov, Barkalov, 2009), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Dolichopus simplex Meigen, 1824 – Volga River basin (Eversmann, 1834), Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Voronezh reg. (Negrobov, 1963), North Caucasus (Negrobov, 1967 c), Murmansk reg. (Negrobov, 1974), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984), Vologda reg. (Grichanov, 2006 b), Kursk reg. (Grichanov, 2007 a), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Kaluga reg. (Grichanov, 2010 b), Dolichopus simulator Parent, 1926 – Primorye (Udovenko, 1970), Dolichopus sinuatus Negrobov et Barkalov, 1978 – Yakutia (Negrobov, Barkalov, 1978), Altai (Negrobov, Barkalov, 2009), Dolichopus smirnovianus Negrobov, 1977 – Primorye (Negrobov, 1977), Dolichopus socer Loew, 1871 – North Ural (Loew, 1871), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Yakutia (Stackelberg, 1933, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Karachay-Cherkessia Republic (Negrobov, Rodionova, 2004 а), Altai (Grichanov, 2007 b, Negrobov, Barkalov, 2009), Dolichopus spinuliformis Maslova, Negrobov, Selivanova, 2012 – Sachalin (Maslova, Negrobov, Selivanova, 2012), Dolichopus spretus Loew, 1871 – Ural (Stackelberg, 1933), Altai (Negrobov, Barkalov, 2009), Dolichopus stackelbergi Smirnov, 1948 – Primorye (Smirnov, 1948 b), Dolichopus subpennatus d’Assis Fonseca, 1976 – Adygea (Grichanov, Volfov, Kustov, 2009), Dolichopus subspretus Negrobov, 1979 Magadan reg. (Negrobov, 1979, Negrobov, Chalaya, 1991), Dolichopus sychevskajae Negrobov et Barkalov, 1978 – Yamal-Nenets autonomous region (Negrobov, Barkalov, 1978), Dolichopus taigensis Smirnov, 1948 Primorye (Smirnov, 1948 b), Yakutia (Negrobov, Chalaya, 1991), Primorye , Magadan reg., Khabarovsk Territory, Kamchatka, Karelia (Negrobov, Maslova, Selivanova, 2011), Dolichopus taimyricus Selivanova, Negrobov et Barkalov, 2012 – Krasnoyarsk Territory (Selivanova, Negrobov, Barkalov, 2012), Dolichopus terminasianae Negrobov, Selivanova, Maslova 2011 Magadan reg. (Negrobov, Selivanova, Maslova, 2011), Dolichopus trangularis Smirnov, 1948 – Primorye (Smirnov, 1948 b), Dolichopus trivialis Haliday, 1832 – Moscow reg. (Fedtchenko A., 1868), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Altai (Grichanov, 2007 b), Adygea (Grichanov, Volfov, Kustov, 2009), Dolichopus tumicosta Negrobov, Grichanov et Barkalov, 2009 – Altai (Negrobov, Grichanov, Barkalov, 2009), Dolichopus tundrensis Barkalov, Negrobov et Grichanov 2009 – Altai (Barkalov, Negrobov, Grichanov, 2009), Dolichopus ukokensis Negrobov et Barkalov, 2009 – Altai (Negrobov, Barkalov, 2009), Dolichopus ungulatus Linnaeus, 1758 – Altai (Motschulsky, 1859, Grichanov, 2007 b), Moscow reg. (Fedtchenko A., 1868, Fedtchenko A., 1891, Fedtchenko B., 1892), Siberia, Altai (Frey, 1915, Stackelberg, 1928 a, 1933), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c, 1968 b, Grichanov, Volfov, Kustov, 2007), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984), Vologda reg. (Grichanov, 2006b), Kursk reg. (Grichanov, 2007 a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Ryazan reg. (Negrobov, Pogonin, 2008), Khanty-Mansi autonomous region (Grichanov, 2010 a), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Dolichopus uniseta Stackelberg, 1929 – Yakutia (Stackelberg, 1929), Primorye (Stackelberg, 1930 b, Stackelberg, 1933, Udovenko, 1970), Far East (Negrobov, Sviridova, 1983), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus uralensis Stackelberg, 1933 – Ural (Stackelberg, 1933, Negrobov, 1970), Dolichopus urbanus Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915, Frey¸ Lundsrom, 1913), Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967d, Grichanov, Volfov, Kustov, 2007), Voronezh reg. (Negrobov, 1972), Altai (Grichanov, 2007 b), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Khanty-Mansi autonomous region (Grichanov, 2010 a), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Dolichopus ussuriensis Stackelberg, 1930 – Primorye (Stackelberg, 1930 b, Stackelberg, 1933, Negrobov, Sviridova, 1983), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus vadimiani Negrobov et Barkalov 1978 Primorye (Negrobov, Barkalov, 1978, Negrobov, Sviridova, 1983), Dolichopus varians Smirnov, 1948 – Primorye (Smirnov, 1948 b), Far East (Negrobov, Sviridova, 1983), Kamchatka (Negrobov, Chalaya, 1991), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus verae Negrobov, 1977 – Kuril Islands (Negrobov, 1977 c), Dolichopus victoris Stackelberg, 1933 – Uzbekistan (Stackelberg, 1933), Altai (Negrobov, Rodionova, 2004 а, Grichanov, 2007 b), Dolichopus violovitshi Negrobov, 1977 – Kuril Islands (Negrobov, 1977 c), Kamchatka (Negrobov, Rodionova, 2004 а), Dolichopus vitripennis Meigen, 1824 – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1921, 1925, 1962), , Moskow reg. (Stackelberg, 1933), Dolichopus wahlbergi Zetterstedt, 1843 – Karelia (Frey, 1915), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Primorye (Stackelberg, 1933), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Voronezh reg. (Negrobov, 1972), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Krasnodar Territory (Grichanov, 2006a), Vologda reg. (Grichanov, Volfov, Kustov, 2006), Dolichopus xanthopyga Stackelberg, 1930 – Primorye (Stackelberg, 1930 b, 1933), Khabarovsk Territory (Grichanov, 2006 a), Dolichopus zernyi Parent, 1927 – Ural (Parent, 1927), Siberia (Stackelberg, 1933), Voronezh reg. (Negrobov, 1965 c, 1966 а, 1972, Negrobov, 1973 c), Astrakhan reg. (Grichanov, 2011), Dolichopus zetterstedti Stenhammer, 1851 – Karelia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Yakutia (Stackelberg, 1933, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Khanty-Mansi autonomous region (Grichanov, 2010 a), Dolichopus zhelochovzevi Negrobov, 1976 – Yakutia (Negrobov, 1976 c, Negrobov, Grichanov, Barkalov, 2009, Grichanov, Bagachanova, 2006).
Epithalassius Mik, 1891.
Epithalassius caucasicus Becker, 1918 – “Black Sea coast of the Caucasus” (Becker, 1918), North Caucasus (Negrobov, Selivanova, 2006).
Guzeriplia Negrobov, 1968.
Guzeriplia chlorina Negrobov, 1968 – North Caucasus (Negrobov, 1968 c), Adygea (Grichanov, Volfov, Kustov, 2009), Krasnodar Territory (Grichanov, 2012 c), Guzeriplia viridana Negrobov, 1978 – North Caucasus (Negrobov, 1978).
Hercostomus Loew, 1857.
Hercostomus aerosus (Fallen, 1823) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko A., 1891, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915, Negrobov, 1974, Grichanov, 2004), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), Primorye (Stackelberg, 1933), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Arkhangelsk reg., Amur reg., Baikal, Mordovia (Chalaya, Negrobov 1989), Sakhalin (Negrobov, Rodionova, 2004 b), Vologda reg. (Grichanov, 2006b), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Hercostomus albibarbus Negrobov, 1976 (Mongolia) – Altai (Grichanov, 2007 b), Hercostomus angustifrons (Staeger 1842) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Voronezh reg. (Negrobov, 1967 a), Ryazan reg. (Negrobov, Pogonin, 1984, 2008), Lipetsk reg., Ryazan reg., Ural (Chalaya, Negrobov 1989), Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Russia (Parent, 1938), North Caucasus (Negrobov, 1967 c), Ryazan reg., Hercostomus anomalipennis Stackelberg, 1933 (Uzbekistan) – Orenburg reg. (Chalaya, Negrobov 1989), Hercostomus arcticus Yang, 1996 (China) – Khabarovsk Territory (Grichanov, 2006 a), Hercostomus armenorum Stackelberg, 1934 (Armenia) – Adygea (Grichanov, Kustov, Volfov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2007), Hercostomus assimilis (Staeger, 1842) – Leningrad reg. (Stackelberg, 1962), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Moscow reg, Ryazan reg., Voronezh reg. (Chalaya, Negrobov 1989), Republic Karachay-Cherkessia (Negrobov, Rodionova, 2004 b), Hercostomus baicalensis Negrobov, 1977 – Yamal-Nenets autonomous region, Buryatia (Negrobov, 1977 a), Hercostomus brevicornis (Staeger, 1842) – Siberia (Frey, 1915), Primorye (Stackelberg, 1934), Russia (Parent, 1938), Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1963, 1967 a), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Moscow reg., Ural (Chalaya, Negrobov 1989), Far East (Negrobov, Rodionova, 2004 b), Murmansk reg. (Grichanov, 2004), Khabarovsk Territory (Grichanov, 2006 a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Hercostomus caucasicus Stackelberg, 1934 – North Caucasus (Stackelberg, 1934, Negrobov, 1965 а, 1967 c, 1968 b, 1970, Grichanov, Volfov, Kustov, 2006, 2007), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Adygea (Grichanov, Volfov, Kustov, 2009), Hercostomus caudatus (Loew, 1859) – Krasnodar Territory (Negrobov, 1965 b), North Caucasus (Grichanov, Volfov, Kustov, 2006), Adygea (Grichanov, Volfov, Kustov, 2009), Hercostomus celer (Meigen, 1824) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Siberia (Frey, 1915), Novgorod reg. (Stackelberg, 1919), Voronezh reg. (Negrobov, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c), Mordovia (Negrobov, Gоlubcova, 1983), Mordovia, Altai (Chalaya, Negrobov 1989), Ryazan reg. (Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Hercostomus chaerophylli (Meigen, 1824) – Krasnodar Territory (Negrobov, 1968 b), North Caucasus (Chalaya, Negrobov 1989), Hercostomus chalybeus (Wiedemann, 1817) – Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1972), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Hercostomus chetifer (Walker, 1849) – Krasnodar Territory (Negrobov, 1965 b, Negrobov, Duhanina, 1984), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Moscow reg. (Chalaya, Negrobov 1989), Hercostomus chrysozygos (Wiedemann, 1817) – Volga River basin (Eversmann, 1834), Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Khabarobsk territory (Negrobov, Rodionova, 2004 b), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Hercostomus conformis (Loew, 1857) – Russia (Parent, 1938), Adygea (Grichanov, Volfov, Kustov, 2009), Krasnodar territory (Grichanov, 2012 c), Hercostomus convergens (Loew, 1857) – Voronezh reg. (Negrobov, 1965 b, 1965 с, 1965, 1966 а, 1972), North Caucasus (Grichanov, Volfov, Kustov, 2006), Hercostomus daubichensis Stackelberg, 1934 – Primorye (Stackelberg, 1934), Far East (Negrobov, Rodionova, 2004 b), Hercostomus dichromopyga Stackelberg, 1934 – Primorye (Stackelberg, 1934), Hercostomus exarticulatus (Loew 1857) – Voronezh reg. (Negrobov, Rodionova, 2004 b), Hercostomus flaveolus Negrobov et Chalaya, 1987 – Primorye (Negrobov, Chalaya, 1987), Hercostomus flavicoxus Negrobov et Logvinovskij, 1977 – Yakutia (Negrobov, Logvinovskii, 1976, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Hercostomus fugax (Loew, 1857) – Russia (Lundstrom, Frey, 1913), Arkhangelsk reg. (Stackelberg, 1934), Russia (Parent, 1938), North Caucasus (Negrobov, 1967 c, 1968 b, Grichanov, Volfov, Kustov, 2006, 2007), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Altai (Grichanov, 2007b), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Hercostomus fuscipennis (Meigen, 1824) – Russia (Parent, 1938), Khabarobsk territory (Negrobov, Rodionova, 2004 b), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Rostov reg. (Grichanov, 2012 c), Hercostomus germanus (Wiedemann, 1817) – Volga River basin (Eversmann, 1834), Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Siberia (Frey, 1915), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962, Stackelberg, 1934), Buryatia, Yakutsk (Negrobov, Rodionova, 2004 b), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Hercostomus grandicercus Negrobov et Nechay, 2009 – Adygea (Negrobov, Nechaj, 2009), Hercostomus inornatus (Loew, 1857) – Russia (Parent, 1938), Hercostomus kedrovicus Negrobov et Logvinovskij 1977 – Primorye (Negrobov, Logvinovskii, 1977, Negrobov, Nechaj, 2011), Hercostomus longiventris (Loew 1857) – North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Hercostomus metallicus (Stannius, 1831) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962, Stackelberg, 1934), Voronezh reg. (Negrobov, 1967 a), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Hercostomus minutus Negrobov et Logvinovskij, 1977 – Primorye (Negrobov, Logvinovskii, 1977), Hercostomus nemorum Smirnov et Negrobov 1977 – Primorye (Smirnov, Negrobov, 1977), Hercostomus nigrilamellatus (Macquart, 1827) – North Caucasus (Negrobov, Rodionova, 2004b), Hercostomus nigripennis (Fallen, 1823) – Ural (Eversmann, 1834), Hercostomus nigriplantis (Stannius, 1831) – Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Mordovia (Negrobov, Gоlubcova, 1983), Kursk reg. (Grichanov, 2007 a), Ryazan reg. (Negrobov, Pogonin, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Hercostomus parvilamellatus (Macquart, 1827) – Russia (Parent, 1938), Hercostomus praeceps Loew, 1869 – Leningrad reg. (Stackelberg, 1962), Burjatia (Negrobov, Rodionova, 2004b), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Hercostomus pseudoceler Stackelberg, 1954 – Primorye (Stackelberg, 1934), Kuril Isl. (Negrobov, Rodionova, 2004b), Hercostomus pterostichoides Stackelberg, 1934 – Primorye (Stackelberg, 1934), Hercostomus radialis Stackelberg, 1934 – Primorye (Stackelberg, 1934), Hercostomus rivulorum Stackelberg, 1934 – Primorye (Stackelberg, 1934), Hercostomus rohdendorfi Stackelberg, 1934 – Primorye (Stackelberg, 1934), Yakutia (Chalaya, Negrobov 1989), Far East (Negrobov, Rodionova, 2004 b), Khabarovsk Territory (Grichanov, 2006 a), Hercostomus rothi (Zetterstedt, 1859) – Vologda reg. (Grichanov, 2006b), Hercostomus rusticus (Meigen, 1824) – Primorye (Stackelberg, 1934), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), Kursk reg., Ural, Krasnoyarsk Territory (Chalaya, Negrobov 1989, Pogonin, Negrobov, 2008), Leningrad reg., Sayan Mountains, Amur. reg. (Negrobov, Rodionova, 2004b), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Yakutia (Grichanov, Bagachanova, 2006), Altai (Grichanov, 2007b), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Dagestan (Grichanov, 2012 c), Hercostomus sahlbergi (Zetterstedt, 1838) – Murmansk reg. (Frey, 1915, Negrobov, 1974), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Primorye (Negrobov, 1968 b), Bashkiria (Chalaya, Negrobov 1989), Ural (Grichanov, 1998), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Adygea (Grichanov, Volfov, Kustov, 2006, 2009), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Dagestan (Grichanov, 2012), Hercostomus stroblianus Becker, 1917 – Russia (Becker, 1917), Krasnodar Territory (Negrobov, 1991), Grichanov, Kustov, Volfov, 2006), Hercostomus sviridovae Negrobov et Tshalaja 1987 – Primorye (Negrobov, Chalaya, 1987), Hercostomus udeorum Stackelberg, 1934 – Primorye (Stackelberg, 1934), Altai (Chalaya, Negrobov, 1989), Amur reg. (Grichanov, 1998), Hercostomus udovenkovae Negrobov et Logvinovskij, 1977 – Primorye (Negrobov, Logvinovskii, 1977), Hercostomus ussurianus Stackelberg 1934 – Primorye (Stackelberg, 1934), Khabarovsk Territory (Grichanov, 1998), Far East (Negrobov, Rodionova, 2004 b), Khabarovsk Territory (Grichanov, 2006 a), Hercostomus varicoloris Becker, 1917 – "Kaukasus, Ostkuste des Schwarzen Meeres" (Becker 1917), Stavropol reg. (Stackelberg, 1941), North Caucasus (Negrobov, 1965 а, 1967 c, 1970, Grichanov, Volfov, Kustov, 2006, 2007), Adygea (Grichanov, Volfov, Kustov, 2009, Grichanov, 2012 c), Hercostomus vivax (Loew, 1857) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Kursk reg., Altai, Ural, Yakutia (Chalaya, Negrobov, 1989), Yakutia (Negrobov, Chalaya, 1991), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2006), Altai (Grichanov, 2007 b), Hercostomus zieheni Parent, 1929 – Amur reg. (Parent, 1929, Selivanova, Negrobov, Maslov, 2008).
Hydrophorus Fallen, 1823.
Hydrophorus albiceps Frey, 1915 – Murmansk reg. (Frey, 1915, Negrobov, 1974), Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Kola Peninsula (Negrobov, 1975 c), Siberia (Negrobov, 1977 a), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Hydrophorus alpinus Wahlberg, 1844 – Archangelsk reg. (Lundstrom, Frey, 1913, Negrobov, 1977 a), Yakutia, mouth of Lena river (Frey, 1915), Murmansk reg. (Negrobov, 1974), Ural (Becker, 1915, 1923), Kola Peninsula (Negrobov, 1975 c, Negrobov, 1975 b), Chukchi AD (Negrobov, Chalaya, 1991), Yakutia (Grichanov, Bagachanova, 2006), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Murmansk reg., Leningrad reg., Arkhangelsk reg., Nenetski, Chukchi and Yamal-Nenets autonomous region (Maslova, Negrobov, Selivanova, 2012), Hydrophorus arcticus Negrobov, 1977 – Chukchi autonomous region (Negrobov, 1977 a, Negrobov, Chalaya, 1991), Yakutia (Grichanov, Bagachanova, 2006), Hydrophorus baicalensis Negrobov, 1977 – Baikal (Negrobov, 1977 е), Hydrophorus balticus (Meigen, 1824) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Baikal (Negrobov, 1977 a), Krasnodar Territory (Negrobov, Duhanina, 1984, Grichanov, Kustov, Volfov, 2006), Yakutia (Grichanov, Bagachanova, 2006), Adygea (Grichanov, Volfov, Kustov, 2009), Hydrophorus bipunctatus (Lehmann, 1822) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Siberia (Frey, 1915), Leningrad reg. (Porchinskij, 1874, Stackelberg, 1921, 1925, 1962,), Voronezh reg. (Negrobov, 1963, 1967 a), Moscow reg., Leningrad reg., Voronezh reg., Buryatia (Negrobov, 1975 c), Ural (Negrobov, 1977b), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Yakutia (Grichanov, Bagachanova, 2006), Kursk reg. (Grichanov, 2007 a), Hydrophorus borealis Loew, 1857 – Leningrad reg. (Stackelberg, 1925, 1962), Arkhangelsk reg., Sverdlovsk reg. (Negrobov, 1975 с), Ural (Negrobov, 1977 a), Murmansk reg. (Grichanov, 2004), Hydrophorus brunneifacies Negrobov, 1977 – Altai (Negrobov, 1977 b, Grichanov, 2007 b), Hydrophorus brunnicosus Loew, 1857 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1966 а, 1967 a), Yaroslav reg., Moscow reg., Ryazan reg., Voronezh reg., Ural, Krasnoyarsk Territory, Yakutia (Negrobov, 1975 c), Ural (Negrobov, 1977 a), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Hydrophorus callostomus Loew, 1857 – Leningrad reg., Yaroslavl reg., Voronezh reg., North Caucasus, Omsk reg., Transbaikalia (Negrobov, 1975 c), Ural (Negrobov, 1977 a), Yakutia (Grichanov, Bagachanova, 2006), Hydrophorus cinipunctus Negrobov, 1975 – Kamchatka (Negrobov, 1975 a), Buryatia (Negrobov, 1975 c), Ural (Negrobov, 1977 a), Magadan reg. (Negrobov, Chalaya, 1991), Hydrophorus femoratus Parent, 1930 – Kamchatka (Parent, 1930), Siberia (Negrobov, 1975 c, Negrobov, 1977 a), Hydrophorus freyi Stora, 1954 – Buryatia, Khabarovsk Territory, Primorye (Negrobov, 1975 c), Siberia (Negrobov, 1977 a), Hydrophorus geminus Frey, 1915 – Murmansk reg. (Frey, 1915, Negrobov, 1974), Siberia (Negrobov, 1977 a), Hydrophorus irinae Negrobov, 1977 – Sverdlov reg. (Negrobov, 1977 g), Kamchatka (Negrobov, Chalaya, 1991), Hydrophorus litoreus Fallén, 1823 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1966 а, 1967 a), Murmansk reg. (Negrobov, 1974), Leningrad reg., Voronezh reg., Tyumen reg., Yakutia (Negrobov, 1975 c), Siberia (Negrobov, 1977 a), Mordovia (Negrobov, Gоlubcova, 1983), Krasnodar Territory (Negrobov, Duhanina, 1984), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Kamchatka (Negrobov, Chalaya, 1991), Yakutia (Grichanov, Bagachanova, 2006), Hydrophorus nebulosus Fallen, 1823 – Siberia (Frey, 1915, Negrobov, 1977 a), Leningrad reg. (Stackelberg, 1925, 1962, Negrobov, 1975 c), Hydrophorus nigrihalteratus Parent, 1930 – Kamchatka (Parent, 1930, Negrobov, 1975 c, Negrobov, 1977 a), Magadan reg. (Negrobov, Chalaya, 1991), Hydrophorus norvegicus Ringdahl, 1928 – Kola Peninsula (Negrobov, 1975 c), Murmansk reg. (Negrobov, 1977 a), Hydrophorus pectinatus Gerstäcker, 1864
– Siberia (Frey, 1915, Negrobov, 1977 a), Leningrad reg. (Stackelberg, 1925, 1962), Tumen reg., Nenets autonomous region (Negrobov, 1975 c), North Caucasus (Grichanov, Volfov, Kustov, 2006), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Hydrophorus pilipes Frey, 1915 – Murmansk reg. (Frey, 1915), Arkhangelsk reg. (Negrobov, 1977 a), Hydrophorus ponojensis Frey, 1915 – Murmansk reg. (Frey, 1915), Hydrophorus praecox (Lehmann, 1822) – Murmansk reg. (Frey, 1915, Negrobov, 1974), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Astrakhan reg., Khabarovsk Territory (Negrobov, 1975 c), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Astrakhan reg. (Grichanov, 2011), Hydrophorus rasnitsyni Negrobov, 1977 – Transbaikalia (Negrobov, 1977 е), Hydrophorus rogenhoferi Mik, 1874 – Siberia (Becker, 1915), Hydrophorus rufibarbis Gerstäcker, 1864 – Murmansk reg. (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Kola Peninsula (Negrobov, 1975 c), Arkhangelsk reg. (Negrobov, 1977 a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Hydrophorus signifer Coquillett, 1899 (synonyms – Hydrophorus magnicornis Frey, 1915, Hydrophorus kolensis Parent, 1934) – Murmansk reg. (Frey, 1915, Negrobov, 1974), Kola Peninsula, Ural (Negrobov, 1975 c, Negrobov, 1977 a), Magadan reg. (Negrobov, Chalaya, 1991), Yakutia (Grichanov, Bagachanova, 2006), Hydrophorus starcus Negrobov et Golubtzov, 2005 – Magadan reg. (Negrobov, Golubtzov 2005), Hydrophorus thibetanus Becker, 1917 – Russia (Becker, 1917), Buryatia (Negrobov, 1977 a), Kamchatka (Negrobov, Chalaya, 1991), Hydrophorus viridis (Meigen, 1824) – Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1972), Leningrad reg., Voronezh reg., Rostov reg., Tumen reg. (Negrobov, 1975 c, Negrobov, 1977 a), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Magadan reg. (Negrobov, Chalaya, 1991), Far East (Negrobov, Golubtzov, 2006), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Krasnodar Territory (Grichanov, 2012 c), Hydrophorus wahlgreni Frey, 1915 – Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1972), Leningrad обл, Kamchatka (Negrobov, 1975 c), Mordovia (Negrobov, Gоlubcova, 1983), Murmansk reg. (Negrobov, 1977 a), Kamchatka (Negrobov, Chalaya, 1991).
Hypophyllus Haliday, 1832.
Hypophyllus crinipes (Staeger, 1842) – Voronezh reg. (Negrobov, 1965 c, 1966 а, 1972), Hypophyllus discipes (Ahrens, 1817) – Russia (Parent, 1938), Hypophyllus obscurellus Fallen, 1823 – Krasnodar Territory (Negrobov, 1965 b), North Caucasus (Negrobov, 1967 c, Grichanov, 1998).
Lamprochromus Mik, 1878
Lamprochromus bifasciatus (Macquart 1827) (synonym – Lamprochromus elegans (Meigen, 1930) – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Lamprochromus buchtojarovi Negrobov Chalaya, 1987 – Chuvashia (Negrobov, Chalaya, 1988), Lamprochrosus speciosus Loew, 1871 – North Caucasus (Grichanov, Volfov, Kustov, 2007), Lamprochromus strobli (Parent, 1924) – Russia (Parent, 1938).
Liancalus Loew, 1857
Liancalus virens (Scopoli, 1763) – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c).
Ludovicius Rondani, 1843.
Ludovicius transcaucasicus Stackelberg, 1941 – North Caucasus (Stackelberg, 1941, Negrobov, 1965 а, 1967 c, 1966).
Machaerium Haliday, 1832.
Machaerium maritimae Haliday, 1832 – Russia (Parent, 1938).
Medetera Fischer von Waldheim, 1819.
Medetera abstrusa Thuneberg, 1955 – Voronezh reg. (Negrobov, 1966 а), Leningrad reg., Voronezh reg., Krasnodar Territory, Novosibirsk reg. (Negrobov, 1971 a), Siberia (Bogdanova, 1975), Mordovia (Negrobov, Gоlubcova, 1983), Medetera acanthura Negrobov & Thuneberg, 1970 – Leningrad reg., Perm reg. (Negrobov, Thuneberg, 1970), Murmansk reg. (Negrobov, Stackelberg, 1972), Medetera acuta Negrobov, 1966 – Leningrad reg. (Negrobov, 1966), Medetera adjaniae Gosseries 1988 (synonym
– Medetera breviseta Parent, 1927) – Perm reg. (Negrobov, Stackelberg, 1972), Arkhangelsk reg. (Negrobov, 1971 a), Medetera alexandri Negrobov, 1979 – Kuril Islands (Negrobov, 1979 a), Medetera ambigua (Zetterstedt, 1843) – Russia (Kowarz, 1877), Krasnoyarsk reg. (Shirskaia, 1961), Siberia (Bogdanova, 1975), Leningrad reg. (Stackelberg, 1925, 1962, Gusev, 1928), Voronezh reg. (Negrobov, 1965 c, 1967 a), Primorye (Negrobov, Stackelberg, 1972), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Medetera apicalis (Zetterstedt, 1843) –Murmansk reg. (Frey, 1915), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962, Gusev, 1928, Zinovev, 1957), North Caucasus (Negrobov, Stackelberg, 1972), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Vologda reg. (Grichanov, 2006 b), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera asiatica Negrobov et Zaitzev, 1979 – Tuva (Negrobov et Zaitzev 1979), Medetera baicalica Negrobov, 1972 – Baikal (Negrobov, Stackelberg, 1972, Negrobov, 1972), Medetera bargusinica Negrobov, 1972 – Buryatia (Negrobov, Stackelberg, 1972), Yakutia (Grichanov, Bagachanova, 2006), Medetera belgica Parent, 1936 – Murmansk reg. (Negrobov, 1971 a, Negrobov, Stackelberg, 1972, Negrobov, 1974), Medetera betulae Ringdahl, 1949 – Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Medetera bidentata Negrobov et Golubtzov, 1991 – Primorye (Negrobov, Gоlubcov, 1993), Medetera bilineata Frey, 1915 – Murmansk reg. (Frey, 1915, Frey¸ Lundsrom, 1913), Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1963, 1967 a), Murmansk reg. (Negrobov, Stackelberg, 1972), Mordovia (Negrobov, Gоlubcova, 1983), Yakutia (Negrobov, Chalaya, 1991), Medetera bisecta Negrobov, 1967 – North Caucasus, Voronezh reg. (Negrobov, 1967 b, Negrobov, Stackelberg, 1972), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera bispinosa Negrobov, 1967 – Voronezh reg. (Negrobov, 1967 b), North Caucasus (Negrobov, Stackelberg, 1972), Medetera borealis Thuneberg, 1955 – Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1972), Primorye (Negrobov, Stackelberg, 1972), Murmansk reg. (Negrobov, 1974), Yakutia (Negrobov, Chalaya, 1991), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera collarti Negrobov, 1967 – North Caucasus (Negrobov, 1967 b), Medetera capillata Negrobov, 1972 – Primorye (Negrobov, Stackalberg, 1972), Medetera complicata Negrobov, 1967
– Perm reg. (Negrobov, 1967 b, Negrobov, Stackelberg, 1972), Medetera delita Negrobov, 1972 – Primorye (Negrobov, Stackelberg, 1972), Medetera diadema (Linnaeus, 1767) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962, Porchinskij, 1874), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c), Ryazan reg. (Negrobov, Pogonin, 1984, 2008), Adygea (Grichanov, Volfov, Kustov, 2006, 2009), Altai (Grichanov, 2007 b), Kursk reg. (Grichanov, 2007 a), North Ossetia (Grichanov, 2012 c), Medetera dichrocera Kowarz, 1877 – Leningrad reg. (Stackelberg, 1962, Zinovev, 1957), Arkhangelsk reg. (Prjahina, Ogibin, 1970, Negrobov, Stackelberg, 1972), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera educata Negrobov, 1979 – Primorye (Negrobov, 1979 b), Medetera excellens Frey, 1909- Leningrad reg. (Stackelberg, 1962, Zinovev, 1957, Negrobov, 1967 b), Primorye (Negrobov, Stackelberg, 1972, 1974 b), Medetera fasciata Frey, 1915 – Leningrad reg. (Zinovev, 1957, Negrobov, Stackelberg, 1972), Medetera fascinator Negrobov, 1972 – Ussuriisk reg. (Negrobov, Stackelberg, 1972), Medetera feminina Negrobov, 1967 – Voronezh reg., Rostov reg., Novgorod reg., North Caucasus (Negrobov, 1967 b), Voronezh reg. (Negrobov, 1972, Negrobov, Stackelberg, 1972), Medetera fissa Negrobov, 1972 – Ussuriisk reg. (Negrobov et Stackelberg, 1972), Medetera flavipes Meigen, 1824 – North Caucasus (Grichanov, Volfov, Kustov, 2006), Medetera freyi Thuneberg, 1955 – Leningrad reg. (Stackelberg, 1962, Negrobov, Stackelberg, 1972), Medetera fumida Negrobov, 1967 – Leningrad reg. (Negrobov, 1967 b), Arkhangelsk reg. (Prjahina, Ogibin, 1970, Negrobov, Stackelberg, 1972), Medetera glaucella Kowarz, 1877 – Russia (Parent, 1938), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Medetera gracilicauda Parent, 1927 – North Caucasus (Grichanov, Volfov, Kustov, 2007), Medetera hymera Negrobov, 1974 – Amur reg. (Negrobov, 1972, Negrobov, Stackelberg, 1974 а), Medetera impigra Collin, 1941 –Novosibirsk reg., Sayan Mountains (Negrobov, 1971 a), Krasnodar Territory (Negrobov, Stackelberg, 1974 а), Siberia (Bogdanova, 1975), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), North Caucasus (Grichanov, Volfov, Kustov, 2007), Medetera incanus Negrobov, 1967 – Leningrad reg. (Negrobov, 1967 b), Medetera incisa Negrobov, 1974 – Primorye (Negrobov, Stackelberg, 1974 a), Medetera incrassatа Frey, 1909 – Leningrad reg. (Stackelberg, 1925, 1962, Negrobov, Stackelberg, 1974 a), Medetera infumata Loew, 1857 – Moscow reg. (Nikitjuk, 1951, 1957), Siberia (Frey, 1915, Tarasova, 1968, Kolomiec, Bogdanova, 1973, Bogdanova, 1974, 1975), Russia (Kowarz, 1877, Frey, 1918), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962, Gusev, 1928, Zinovev, 1957), Krasnoyarsk reg. (Shirskaia, 1961), Voronezh reg. (Negrobov, 1965 c), Primorye (Negrobov, Stackelberg, 1974 а), Ryazan reg. (Negrobov, Pogonin, 1984), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera infuscatа Negrobov, 1974 – Primorye (Negrobov, Stackelberg, 1974 a), Medetera inspissata Collin, 1952 – Krasnodar Territory (Negrobov, Stackelberg, 1974 а), Medeterajacula (Fallén, 1823) – Moscow reg. (Fedtchenko A., 1968, Fedtchenko B., 1892), Siberia (Frey, 1915), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1921, 1925, 1962, Gusev, 1928, Zinovev, 1957), Voronezh reg. (1963, 1965 c, 1966 а, 1967 a), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Sayan Mountains (Negrobov, 1985, Negrobov, Selivanova, Maslova, 2010), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Vologda reg. (Grichanov, 2006 b), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Altai (Grichanov, 2007 b), Kursk reg. (Grichanov, 2007 a), Kaluga reg. (Grichanov, 2010 b), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera jacuta Negrobov, 1972 – Yakutia (Negrobov, Stackelberg, 1974 а, Grichanov, Bagachanova, 2006), Medetera jugalis Collin, 1941 – Baikal (Negrobov, 1971 a, Negrobov, Stackelberg, 1974 а), Medetera kerzhneri Negrobov 1966 (Kasachstan) – Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera krivosheinae Negrobov, 1968 – Yaroslav reg. (Negrobov, 1968 c, Negrobov, Stackelberg, 1974 а), Medetera lamprostomoides Negrobov, 1974 – Yakutia (Negrobov, Stackelberg, 1974 а, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Medetera longicauda Becker, 1917 – Russia (Becker, 1917), Voronezh reg. (Negrobov, 1971 a, 1972, Negrobov, Stackelberg, 1974 а, Negrobov, 1977 g), Medetera lorea Negrobov, 1967 – Voronezh reg. (Negrobov, 1967 b, Negrobov, Stackelberg, 1974 а), Medetera melancholica Lundbeck, 1912 – Arkhangelsk reg. (Prjahina, Ogibin, 1970), Siberia (Tarasova, 1965, 1968, Kolominets, Bogdanova, 1973, Bogdanova, 1974, 1975), Leningrad reg. (Stackelberg, 1925, 1962, Zinovev, 1957), Murmansk reg. (Negrobov, 1974, Negrobov, Stackelberg, 1974 а), Medetera meridionalis Negrobov, 1967 - Voronezh reg., Rostov reg., Krasnodar territory, Volograd reg., Penza reg., Adygea, Orenburg reg., Altai (Negrobov, 1967 b, 1972), East Siberia (Negrobov, Stackelberg, 1974 а), Medetera micacea Loew, 1857 – Russia (Kowarz, 1877), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1965 c, 1967 a), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Adygea (Grichanov, Volfov, Kustov, 2009), Medetera mixta Negrobov, 1967 – Lipetsk reg., Voronezh reg., Bashkiria, North Caucasus (Negrobov, 1968 a, Grichanov, Volfov, Kustov, 2006), Voronezh reg. (Negrobov, 1972), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Medetera mucronata Negrobov et Golubtzov, 1991 – Primorye (Negrobov, Gоlubcov, 1993), Medetera muralis Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Russia (Parent, 1938), North Caucasus (Negrobov, 1967 b, Negrobov, Stackelberg, 1974а, Grichanov, Volfov, Kustov, 2007), Krasnodar Territory (Negrobov, Stackelberg, 1974 b, Grichanov, 2012 c), Adygea (Grichanov, Volfov, Kustov, 2009), Medetera murina Becker, 1917 – Russia (Becker, 1917), North Caucasus (Negrobov, Stackelberg, 1974 а), Medetera nebulosa Negrobov, 1974 – Primorye (Negrobov, Stackelberg, 1974 а), Medetera nitida (Macquart, 1834) – Arkhangelsk reg. (Prjahina, Ogibin, 1970), Leningrad reg. (Stackelberg, 1925, 1962, Zinovev, 1957), Primorye , Ural (Negrobov, Stackelberg, 1974 а), Medetera obscura (Zetterstedt, 1838) – Siberia (Frey, 1915, Negrobov, Stackelberg, 1974 а, Bogdanova, 1974, 1975), Leningrad reg. (Stackelberg, 1925, 1962, Zinovev, 1957), Murmansk reg. (Negrobov, 1974), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera pallens Negrobov, 1967 – Krasnodar Territory, North Caucasus (Negrobov, 1967 b), North Caucasus (Negrobov, Stackelberg, 1974 b), Adygea (Grichanov, Kustov, Volfov, 2006), Medetera pallidior (Stackelberg, 1937) – Turkmenia (Stackelberg, 1937), North Caucasus (Grichanov, Volfov, Kustov, 2007), Medetera pallipes (Zetterstedt, 1843) – Russia (Kowarz, 1877), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962, Gusev, 1928, Zinovev, 1957), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, Stackelberg, 1974 b), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Krasnodar Territory (Grichanov, Volfov, Kustov, 2006), Stavropol Territory (Grichanov, Volfov, Kustov, 2007), Adygea (Grichanov, Volfov, Kustov, 2009), Medetera parenti Stackelberg, 1925 – Leningrad reg. (Stackelberg, 1925, 1962), Baikal (Parent, 1927), Voronezh reg. (Negrobov, 1967 a), North Caucasus (Negrobov, Stackelberg, 1974 b), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Medetera peloria Negrobov, 1967 –North Caucasus (Negrobov, 1967), Medetera penicillata Negrobov, 1970 – Primorye (Negrobov, 1970 a), Medetera perfida Parent, 1932 – North Caucasus (Negrobov, Stackelberg, 1974 b), Adygea (Grichanov, Volfov, Kustov, 2009), Medetera petrophiloides Parent, 1925– Rostov обл, Voronezh reg. (Negrobov, 1965 b), Voronezh reg. (Negrobov, 1972, Negrobov, Stackelberg, 1974 b), Krasnodar Territory (Negrobov, Duhanina, 1984), Medetera pinicola Kowarz, 1877
– Leningrad reg. (Stackelberg, 1925, 1962, Gusev, 1928, Zinovev, 1957), Krasnoyarsk reg. (Shirskaia, 1961), Voronezh reg. (Negrobov, 1963, 1963, 1967 а), Arkhangelsk reg. (Prjahina, Ogibin, 1970), Siberia (Tarasova, 1968, Kolomiec, Bogdanova, 1973, 1974, Negrobov, Stackelberg, 1974 b, Bogdanova, 1975), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera plumbella Meigen, 1824 – Voronezh reg. (Negrobov, 1967 a), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Medetera prjachinae Negrobov, 1974 – Arkhangelsk reg. (Negrobov, Stackelberg, 1974 b), Medetera protuberans Negrobov, 1967 – Leningrad reg. (Negrobov, 1967 b, Negrobov, Stackelberg, 1974 b), Medetera pseudoapicalis Thuneberg, 1955 – Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1972), North Caucasus (Negrobov, Stackelberg, 1974 b), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera ravida Negrobov, 1970 – Primorye (Negrobov, 1970 a), Medetera relicta Negrobov, 1967 – North Caucasus, Lipetsk reg., Voronezh reg., North Caucasus, Adygea (Negrobov, 1967 b), Voronezh reg. (Negrobov, 1972), Lipetsk reg. (Negrobov, Stackelberg, 1974 b), Krasnodar Territory (Grichanov, 2012 c), Medetera rufipes Negrobov, 1974 – Primorye (Negrobov, Stackelberg, 1974 b), Medetera seguyi sphaeroidea Negrobov, 1967 – North Caucasus (Negrobov, 1967 b, Negrobov, Stackelberg, 1974 b), Adygea (Grichanov, Volfov, Kustov, 2009), Medetera senicula Kowarz, 1877 – Leningrad reg. (Stackelberg, 1925, 1962, Negrobov, Stackelberg, 1974 b), Khanty-Mansi autonomous region (Grichanov, 2010 a), Medetera sibirica Negrobov 1974 Khanty-Mansi autonomous region (Grichanov, 2010 a), Medetera setiventris Thuneberg, 1955 – Novosibirsk reg. (Negrobov, Stackelberg, 1974 b), Leningrad reg. (Stackelberg, 1962), Siberia (Bogdanova, 1975), Medetera signaticornis Loew, 1857 – Arkhangelsk reg. (Frey, 1918, Prjahina, Ogibin, 1970), Murmansk reg. (Frey¸ Lundsrom, 1913), Leningrad reg. (Stackelberg, 1925, 1962, Gusev, 1928, Zinovev, 1957, 1958), Krasnoyarsk reg. (Shirskaia, 1961), Novorossisk reg., Tuva, North Caucasus (Negrobov, 1970 b), Primorye (Negrobov, Stackelberg, 1974 b), Siberia (Bogdanova, 1975), Khanty-Mansi autonomous region (Grichanov, 2010 a), Medetera sphaeropyga Negrobov 1972 – Ussuriisk reg. (Negrobov, 1972), Primorye (Negrobov, Stackelberg, 1974 b), Medetera spinulicauda Negrobov, 1974 – Amur reg. (Negrobov, Stackelberg, 1974 b), Medetera stackelbergiana Negrobov, 1967 – Voronezh reg. (Negrobov, 1967 b), Primorye (Negrobov, Stackelberg, 1974 b), Medetera striata Parent, 1927 – Voronezh reg., Novosibirsk reg., North Caucasus, Murmansk reg. (Negrobov, 1971 a), Voronezh reg. (Negrobov, 1972), Murmansk reg. (Negrobov, Stackelberg, 1974b, Negrobov, 1974), Medetera stylata Negrobov, 1974 – Primorye(Negrobov, Stackelberg, 1974 b), Medetera subtristis Negrobov, 1974 – Novosibirsk reg., Baikal, Buryatia (Negrobov, Stackelberg, 1974 b), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera sutshanica Negrobov, 1972 – Ussuriisk reg. (Negrobov, 1972), Primorye (Negrobov, Stackelberg, 1974 b), Medetera tarasovae Negrobov, 1972 – Novosibirsk reg. (Negrobov, 1972), Novosibirsk reg., Amur reg. (Negrobov, Stackelberg, 1974 b), Medetera tenuicauda Loew, 1857 – Russia (Kowarz, 1877), Voronezh reg. (Negrobov, 1971 a, Negrobov, 1972), Rostov reg. (Negrobov, Stackelberg, 1974 b), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Medetera tristis (Zetterstedt, 1840) – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962, Zinovev, 1957), Primorye (Negrobov, 1977 a), Medetera truncorum Meigen, 1824 – Russia (Kowarz, 1877), North Caucasus (Negrobov, 1967 c, Negrobov, 1977 a, Grichanov, Volfov, Kustov, 2006), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), Yakutia (Grichanov, Bagachanova, 2006, Grichanov, Tomkovich, 2009), Medetera thunebergi Negrobov, 1967 – Leningrad reg, Nizhni Novgorod reg., Orenburg reg., Novosibirsk reg., Amur reg. (Negrobov, 1967), Medetera tuterculosa Negrobov, 1977 – Primorye (Negrobov, 1977 a), Medetera tumidula Negrobov, 1967 – North Caucasus, Adygea (Negrobov, 1967 b), North Caucasus (Negrobov, 1977 a), Medetera ussuriana Negrobov, 1977 – Primorye (Negrobov, 1977 b), Medetera vagans Becker, 1917 – Russia (Becker, 1917), Leningrad reg. (Negrobov, 1971 a), Murmansk reg. (Negrobov, 1974), Primorye (Negrobov, 1977b), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Medetera veles Loew, 1861 – Yakutia (Grichanov, Bagachanova, 2006), Khanty-Mansi autonomous region (Grichanov, 2010 a), Medetera victoris Negrobov 1972 – Ussuriisk reg. (Negrobov, 1972), Primorye (Negrobov, 1977b), Medetera virescens Negrobov, 1965 – Leningrad reg. (Negrobov, 1967 b), Medetera zinovjevi Negrobov, 1967 – Arkhangelsk reg. (Prjahina, Ogibin, 1970), Perm reg. (Negrobov, 1967 b), Primorye (Negrobov, 1977 a).
Melanostolus Kowarz, 1884.
Melanostolus melancholicus (Loew, 1869) – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), North Caucasus (Negrobov, 1967 c), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Melanostolus nigricilius (Loew, 1871) – Russia (Parent, 1938), Yakutia (Grichanov, Bagachanova, 2006), Melanostolus tatianae Negrobov, 1965 – North Caucasus (Negrobov, 1965 a, 1967 c), Krasnodar Territory (Negrobov, 1965 b), Krasnodar Territory, North Caucasus, Adygea (Negrobov, 1965).
Mesorhaga Schiner, 1868.
Mesorhaga dimi Negrobov, 1984 – Kuril Islands (Negrobov, 1984), Mesorhaga pilosa Negrobov, 1979 – Primorye (Negrobov, 1979 b).
Micromorphus Mik, 1878.
Micromorphus albipes albipes (Zetterstedt, 1843) – Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c), Micromorphus albipes claripennis (Strobl, 1899) – Leningrad reg. (Stackelberg, 1962), Micromorphus amurensis Negrobov, 2000 – Amur reg. (Negrobov, 2000), Micromorphus jakutensis Negrobov, 2000
– Yakutia (Negrobov, 2000, Grichanov, Bagachanova, 2006), Micromorphus shamshevi Negrobov, 2000 – North Caucasus (Grichanov, Volfov, Kustov, 2006).
Nematoproctus Loew, 1857.
Nematoproctus daubichensis Stackelberg et Negrobov, 1976 – Primorye (Stackelberg, Negrobov, 1976), Nematoproctus distendens (Meigen, 1824) – Russia (Parent, 1938), Leningrad reg. (Stackelberg, 1962, Stackelberg, Negrobov, 1976), Nematoproctus longifilus Loew, 1857 – Russia (Parent, 1938), Voronezh reg. (Negrobov, 1977 g), Nematoproctus praesectus Loew, 1869 – Leningrad reg. (Stackelberg, 1925, 1962, Stackelberg, Negrobov, 1976), Kursk reg. (Grichanov, 2007 a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008).
Nepallomyia Hollis, 1964
Nepallomyia (Neurigonella) tatjanae Negrobov, 1984 – Primorye (Negrobov, 1984).
Neurigona Rondani, 1856.
Neurigona abdominalis (Fallén, 1823) – Leningrad reg. (Stackelberg, 1925, 1962), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), Neurigona anomaloptera Negrobov, 1987 – Kuril Islands (Negrobov, 1987), Neurigona davshinica Negrobov, 1987 – Buryatia (Negrobov, 1987), Neurigona erichsoni (Zetterstedt, 1843)
– Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Krasnodar Territory (Negrobov, Duhanina, 1984, Grichanov, 2012 c), North Caucasus (Grichanov, Volfov, Kustov, 2007), Adygea (Grichanov, Volfov, Kustov, 2009), Neurigona febrilata Negrobov et Fursov, 1988 – Krasnodar Territory (Negrobov, Fursov, 1988), Neurigona flavella Negrobov, 1987 – Primorye (Negrobov, 1987), Neurigona grossa Negrobov, 1987 – Primorye (Negrobov, 1987), Neurigona helva Negrobov et Zurikov, 1990 – Krasnodar Territory, North Caucasus (Negrobov, Tsurikov, 1990), Neurigona kasparyani Negrobov, 1987 – Sakhalin (Negrobov, 1987), Neurigona lineata (Oldenberg, 1904) – Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Neurigona longipes (Becker, 1918) – Russia (Becker, 1918), Neurigona micropyga Negrobov, 1987 – Kuril Islands (Negrobov, 1987), Neurigona ninae Negrobov, 1987 – Primorye (Negrobov, 1987), Neurigona pallida (Fallén, 1823) – Orenburg reg. (Becker, 1917, Wnukowsky, 1932, 1936), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1972), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Khanty-Mansi autonomous region (Grichanov, 2010 a), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Rostov reg. (Grichanov, 2012 c), Neurigona pseudolongipes Negrobov, 1987 – Krasnodar Territory (Negrobov, 1987), Neurigona pullata Negrobov, 1988 – Primorye (Negrobov, Fursov, 1988), Neurigona quadrifasciata (Fabricius, 1781) – Moscow reg. (Fedtchenko A., 1868), Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Neurigona semilata Negrobov et Fursov, 1988 – Krasnodar Territory, North Caucasus (Negrobov, Fursov, 1988), Neurigona subcilipes Negrobov, 1988 – North Caucasus (Negrobov, Fursov, 1988, Grichanov, Volfov, Kustov, 2006), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Adygea (Grichanov, Volfov, Kustov, 2009), Neurigona suturalis (Fallén, 1823) – Leningrad reg. (Stackelberg, 1925, 1962), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Neurigona uralensis (Becker, 1918) – Ural (Becker, 1918), Neurigona verrichterae Negrobov et Fursov, 1988 – North Caucasus (Negrobov, Fursov, 1988, Grichanov, Volfov, Kustov, 2007), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006).
Orthoceratium Schrank, 1803.
Orthoceratium lacustre (Scopoli, 1763) – North Caucasus (Grichanov, Volfov, Kustov, 2007).
Paraclius Bigot, 1859.
Paraclius argenteus Negrobov, 1984 – Primorye (Negrobov, 1984).
Peloropeodes Wheeler, 1890.
Peloropeodes acuticornis (Oldenberg, 1916) – Krasnodar Territory (Negrobov, Duhanina, 1984), North Caucasus (Grichanov, Popov, 2007).
Peodes Loew, 1857.
Peodes forcipatus Loew, 1857 – Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Ural, North Caucasus (Grichanov, 2012), Peodes petsamoensis Frey, 1930 – Russia (Frey, 1930), Murmansk reg. (Grichanov, 2012), Peodes yeniseiensis Grichanov, 2012 – Krasnoyarsk Territory (Grichanov, 2012 d).
Poecilobothrus Mik, 1878.
Poecilobothrus basilicus (Loew, 1869) – North Caucasus (Grichanov, Volfov, Kustov, 2007), Poecilobothrus bigoti Mik, 1883 – Krasnodar Territory (Negrobov, 1965 b), North Caucasus (Negrobov, 1967 c), Poecilobothrus comitialis (Kowarz, 1867) – Russia (Parent, 1938), Voronezh reg. (Negrobov, 1965 c, 1966 а, 1972, Negrobov, 1977 g), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Poecilobothrus ducalis (Loew, 1857) – Vologda reg. (Grichanov, 2006 b), Poecilobothrus fumipennis (Stannius, 1831) – Adygea (Negrobov, 1965 b), North Caucasus (Negrobov, 1967 c), Voronezh reg. (Negrobov, 1977 g), Krasnodar Territory (Negrobov, Korneev, 2010), Poecilobothrus infuscatus (Stannius, 1767) – Krasnodar Territory (Negrobov, Duhanina, 1984), Poecilobothrus regalis (Meigen, 1824) – Russia (Parent, 1938), Voronezh reg. (Negrobov, 1965 c, 1972, Negrobov, Korneev, 2010), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Krasnodar Territory (Grichanov, Volfov, Kustov, 2006).
Pseudoxanthochlorus Negrobov, 1977.
Pseudoxanthochlorus micropygus Negrobov, 1977 – Primorye (Negrobov, 1977е).
Rhaphium Meigen, 1803.
Rhaphium albifrons Zetterstedt, 1843 – Leningrad reg. (Stackelberg, 1962), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Mordovia, Krasnoyarsk Territory, Altai, Irkutsk reg., Khabarovsk Territory, Primorye (Maslova, Negrobov, Selivanova, 2012), Rhaphium albomaculatum Beck, 1891 – North Caucasus (Negrobov, 1965), Rhaphium antennatum Carlier, 1835 – Moscow reg. (Fedtchenko A., 1868), Murmansk reg. (Lundsrom, Frey¸ 1913, Frey, 1915,), Russia (Parent, 1938), Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1972), Mordovia (Negrobov, Gоlubcova, 1983), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2006), Astrakhan reg. (Grichanov, 2011), Rhaphium appendiculatum Zetterstedt, 1849 (synonym – Rhaphium macrocerum Meigen, 1824) – Moscow reg. (Fedtchenko A., 1868), Krasnodar Territory (Negrobov, Duhanina, 1984), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), Rhaphium basale Loew, 1850 – Yakutia (Negrobov, 1976 b, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Rhaphium beringiense Negrobov, 1979 – Altai (Negrobov, Barkalov, Selivanova, 2012), Rhaphium boreale Van Duzee, 1923 – Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Altai (Grichanov, 2007 b), Rhaphium borisovi Negrobov, Barkalov & Selivanova, 2012 – Taimyr (Negrobov, Barkalov, Selivanova, 2012), Rhaphium brooksi Negrobov, Barkalov et Selivanova, 2011 – Magadan reg. (Negrobov, Barkalov, Selivanova, 2011), Rhaphium caliginosum Meigen, 1824 (synonym – Rhaphium zetterstedti Parent, 1925) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915), Russia (Parent, 1918, Parent, 1938), Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c), Voronezh reg. (Negrobov, 1972), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), Adygea (Grichanov, Volfov, Kustov, 2006, 2009), Kursk reg. (Grichanov, 2007 a), Rhaphium commune Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Russia (Lundstrom, Frey, 1913), Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Voronezh reg. (Negrobov, 1972), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Yakutia (Grichanov, Bagachanova, 2006), Rhaphium confine Zetterstedt, 1843 – Murmansk reg. (Frey, 1915), Taimyr (Negrobov, Barkalov, Selivanova, 2012), Rhaphium crassipes Meigen, 1824 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Lundstrom, Frey, 1913, Frey, 1915), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Kamchatka (Parent, 1930), Russia (Parent, 1938), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Yakutia (Negrobov, Chalaya, 1991), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Khanty-Mansi autonomous region (Grichanov, 2010 a), Rhaphium dichromum Negrobov, 1976 – Buryatia (Negrobov, 1976 a), Magadan reg. (Negrobov, Chalaya, 1991), Murmansk reg. (Grichanov, 2004), Altai (Grichanov, 2007 b), Khanty-Mansi autonomous region (Grichanov, 2010 a), Rhaphium discigera Stenhammer, 1850 – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1965 c), North Caucasus (Negrobov, 1967 c), Rhaphium discolor Zetterstedt 1838 (synonym – Rhaphium riparium Meigen, 1824) – Murmansk reg. (Frey, 1915, Negrobov, 1968 b, 1974), Leningrad reg. (Stackelberg, 1962), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Rhaphium dispar Coquillett, 1898 (synonym – Rhaphium argyroides Parent, 1926) – Primorye (Negrobov, 1977), Magadan reg. (Negrobov, Chalaya, 1991), Rhaphium elegantulum Meigen, 1824 – Murmansk reg. (Frey, 1915, Grichanov, 2004), Ural (Becker, 1915), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Kamchatka (Parent, 1930), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1972), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Yakutia (Negrobov, Chalaya, 1991), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Khanty-Mansi autonomous region (Grichanov, 2010 a), Rhaphium essoensis Negrobov, 1979 – Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Rhaphium fasciatum Meigen, 1824 – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Rhaphium fascipes Meigen, 1824 – Moscow reg. (Fedtchenko A., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Krasnoyarsk Territory (Negrobov, Pogonin, 2006, Grichanov, Kustov, Volfov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Rhaphium firsovi Stackelberg et Negrobov, 1976 – Primorye (Negrobov, 1976 a), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Rhaphium flavilabre Negrobov, 1979 – Khabarovsk Territory (Grichanov, 1998), Rhaphium glaciale Ringdahl, 1920 – Krasnoyarsk Territory, Buryatia, Yakutia, Magadan reg. (Negrobov, 1976 b), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Magadan reg. (Grichanov, 1998), Murmansk reg. (Grichanov, 2004), Khanty-Mansi autonomous region (Grichanov, 2010a), Rhaphium gravipes Haliday 1851 (synonym – Rhaphium longilamellatum Kowarz, 1867) – Leningrad reg. (Stackelberg, 1925, 1962), Rhaphium gruniniani Negrobov, 1979 – Altai (Grichanov, 2007 b), Rhaphium gussakovskii Stackelberg et Negrobov, 1976 – Primorye (Negrobov, 1976 a), Rhaphium holmgreni (Mik 1878) (synonym – Rhaphium luteipennis (Frey 1915)) – Siberia (Frey, 1915), Rhaphium intermedium Becker, 1918 – Leningrad reg. (Stackelberg, 1962), Rhaphium jamalensis Negrobov, 1986 – Ямал (Negrobov, 1986 a), Taimyr (Negrobov, Barkalov, Selivanova, 2012), Rhaphium lanceolatum Loew, 1850 – Murmansk reg. (Frey¸ Lundsrom, 1913), Vologda reg. (Grichanov, 2006 b), Rhaphium laticorne Fallén, 1823 (synonym – Rhaphium subnudipes Zetterstedt, 1859) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey¸ Lundsrom, 1913), Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 а), North Caucasus (Negrobov, 1967 c), Murmansk reg. (Negrobov, 1974), Mordovia (Negrobov, Gоlubcova, 1983), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Altai (Grichanov, 2007b), Krasnodar Territory (Grichanov, 2012 c), Rhaphium latifacies Negrobov, 1986 – Yakutia (Negrobov, 1986 a), Rhaphium latimanum Kahanpaa, 2007 – Khanty-Mansi autonomous region (Grichanov, 2010 a), Taimyr (Negrobov, Barkalov, Selivanova, 2012), Rhaphium lehri Negrobov, 1977 – Voronezh reg. (Negrobov, Shamshev, 1982), Rhaphium longicorne (Fallen, 1823) – Murmansk reg. (Frey, 1915), Leningrad reg. (Stackelberg, 1925), Rhaphium macalpini Negrobov, 1986 – Yakutia (Negrobov, 1986 a), Rhaphium micans Meigen, 1824 – Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1967 а), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Krasnodar Territory (Negrobov, Korneev, 2010), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Khabarovsk Territory (Grichanov, 2006 a), Astrakhan reg. (Grichanov, 2011), Rhaphium monotrichum Loew, 1850 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1967 a), Mordovia (Negrobov, Gоlubcova, 1983), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2007), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Rhaphium nasutum Fallén, 1823 – Moscow reg. (Fedtchenko A., 1868), Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1972), Ryazan reg. (Negrobov, Pogonin, 1984), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Rhaphium nigribarbatum Becker, 1900 – Ural (Becker, 1915), Arkhangelsk reg. (Frey, 1918), Murmansk reg. (Frey¸ Lundsrom, 1913, Yakutia (Negrobov, Chalaya, 1991), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Altai (Grichanov, 2007 b), Ryazan reg. (Negrobov, Pogonin, 2008), Rhaphium nudiusculum Negrobov, 1976 – Buryatia (Negrobov, 1976 a), Yakutia (Negrobov, Chalaya, 1991), Rhaphium nuortevai Negrobov, 1977 – Primorye (Negrobov, 1977 d), Jewish autonomous region (Grichanov, 1998), Rhaphium patellitarse Becker, 1900 – Ural (Becker, 1915), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Taimyr (Negrobov, Barkalov, Selivanova, 2012), Rhaphium patulum Raddatz, 1873 – Leningrad reg. (Stackelberg, 1921, 1925, 1962), Murmansk reg. (Grichanov, 2004), Rhaphium pectinatum (Loew, 1859) (synonym – Porphyrops pectinata Loew, 1859) – Voronezh reg. (Negrobov, 1965 c, 1966 а, 1972), North Caucasus (Negrobov, 1967 c), Rhaphium penicillatum Loew, 1850 – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c), Voronezh reg. (Negrobov, 1972), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Rhaphium richterae Negrobov, 1977 – Kuril Islands (Negrobov, 1977 d), Rhaphium riparium (Meigen 1824) (synonym – Rhaphium praerosum Loew, 1850) – Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c), Voronezh reg. (Negrobov, 1972), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Rhaphium rivale Loew, 1869 – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Rhaphium sachalinense Negrobov, 1979 – Yakutia (Negrobov, 1979 a), Rhaphium sibiricus Negrobov, Barkalov et Selivanova, 2011 – Altai (Negrobov, Barkalov et Selivanova, 2011), Rhaphium sichotense Negrobov, 1979 – Krasnoyarsk Territory (Negrobov, Pogonin, 2006), Rhaphium stackelbergi Negrobov, 1976 – Primorye (Negrobov, 1976 a), Rhaphium suavis Loew, 1859 – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c), Voronezh reg. (Negrobov, 1972), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Rhaphium subtridactylum Negrobov, Barkalov et Selivanova, 2011 – Altai (Negrobov, Barkalov et Selivanova, 2011), Rhaphium tibiale (von Roser 1840) (synonym – Rhaphium fractum Loew, 1850) – Leningrad reg. (Stackelberg, 1925, 1962), Rhaphium trifidum Becker, 1918 – Baikal (Negrobov, 1976 c), Rhaphium tridactylum Frey, 1915 – Khabarovsk Territory (Negrobov, 1977 b), Taimyr (Negrobov, Barkalov, Selivanova, 2012), Rhaphium tripartitum Frey, in Lundstrom, Frey, 1913 – Murmansk reg. (Frey¸ Lundsrom, 1913), Kola Peninsula (Negrobov, 1977 b), Taimyr (Negrobov, Barkalov, Selivanova, 2012), Rhaphium umbripenne Frey, 1915 – Murmansk reg. (Frey, 1915), Leningrad reg. (Stackelberg, 1962), Yakutia (Negrobov, Chalaya, 1991), Khanty-Mansi autonomous region (Grichanov, 2010 a), Rhaphium venustum Negrobov, 1977 – Primorye (Negrobov, 1977 d).
Scellus Loew, 1857.
Scellus alactaga Stackelberg, 1951 – Yakutia, Chita reg. (Stackelberg 1951 b), Yakutia (Grichanov, Bagachanova, 2006), Scellus gallicanus Becker, 1909 – Yakutia (Stackelberg 1951 b, Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006), Negrobov, Selivanova, Maslova, 2010), Altai (Grichanov, 2007 b), Scellus notatus (Fabricius, 1781) – Ural (Becker, 1915), Arkhangelsk reg. (Frey, 1918), Russia (Parent, 1938), Siberia (Stackelberg 1951 b), North Caucasus (Grichanov, Volfov, Kustov, 2006), Scellus obuchovae Stackelberg, 1951 –Chkalovsk, Verkhnaya Dneprovka (Stackelberg, 1951), Voronezh reg. (Negrobov, 1963, 1965 c, Negrobov, 1977 g), Scellus spinimanus (Zeterstedt, 1843) – Siberia (Frey, 1915), Ural (Becker, 1915, Becker, 1923), Yakutia (Negrobov, Chalaya, 1991, Grichanov, Bagachanova, 2006).
Sciapus Zeller, 1842.
Sciapus albifrons Meigen, 1830 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1965 c, 1972), Ryazan reg. (Negrobov, Pogonin, 1984), Leningrad reg., Moscow reg., Voronezh reg., Novosibirsk reg. (Negrobov, Selivanova, 1994), Ryazan reg. (Negrobov, Pogonin, 2008), Sciapus contristans Wiedemann, 1817 – Moscow reg. (Fedtchenko A., 1868), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925), Leningrad reg., Novgorod reg., Moscow reg. (Negrobov, Selivanova, 1994), Sciapus flavicinctus Loew, 1857 – Krasnodar Territory (Shamshev, 1990, Grichanov, 2012c), North Caucasus (Grichanov, Volfov, Kustov, 2007), Sciapus frater Parent, 1927 – North Caucasus (Negrobov, 1965 а, 1965 b) Sciapus gracilipes (Loew, 1871) – Voronezh reg. (Negrobov, 1965 b, 1965 c, 1966 а, 1972, Negrobov, Selivanova, 1994), Sciapus incognitus Negrobov et Shamshev, 1996 – Primorye (Negrobov, Shamshev, 1986b), Sciapus lobipes Meigen, 1824 – Leningrad reg. (Stackelberg, 1925, 1962), Leningrad reg., Moscow reg. (Negrobov, Selivanova, 1994), Sciapus longulus Fallén, 1823 – Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1963, 1972), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Leningrad reg., Moscow reg., Lipetsk reg., Voronezh reg. (Negrobov, Selivanova, 1994), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Kursk reg. (Grichanov, 2007 a), Sciapus maritimus Becker, 1918 – Leningrad reg. (Stackelberg. 1962), North Caucasus (Negrobov, 1967 c), Leningrad reg., North Caucasus, Krasnodar Territory (Negrobov, Selivanova, 1994), Sciopus nervosus Lehmann, 1822 – Leningrad reg. (Stackelberg, 1925, 1962), Leningrad reg., Moscow reg., Ural, Primorye (Negrobov, Selivanova, 1994), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Irkutsk reg. (Maslova, Negrobov, Selivanova, 2010), Sciapus paradoxus paradoxus Negrobov et Shamshev, 1986 – Primorye (Negrobov, Shamshev, 1986 b), Sciopus paradoxus sachalinensis Negrobov et Shamshev, 1986 – Yakutia (Negrobov, Shamshev, 1986 b), Sciapus platypterus Fabricius, 1805 – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1965 c, 1967 а, Negrobov, Selivanova, 1994), Ryazan reg. (Negrobov, Pogonin, 1984), Leningrad reg., Moscow reg., Ryazan reg. (Negrobov, Pogonin, 2008), Tatarstan (Negrobov, Korneev, Selivanova, 2010), Sciapus polozhentsevi Negrobov, 1977 – North Caucasus (Negrobov, 1977 f), North Caucasus, Krasnodar Territory, Adygea (Negrobov, Selivanova, 1994), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), Sciapus richterae Negrobov et Grichanov, 2010 – Krasnodar Territory (Grichanov, 2012 c), Sciapus roderi Parent, 1930 – Primorye (Parent, 1929), Amur reg. (Negrobov, Selivanova, 1994), Sciapus sibiricus Negrobov et Shamshev, 1986 – Sayan Mountains (Negrobov, Shamshev, 1986 b), Sciapus spiniger Zetterstedt, 1859 – Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), Sciapus wiedemanni Fallén, 1823 – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 с, 1967 a), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Leningrad reg., Voronezh reg. (Negrobov, Selivanova, 1994).
Setihercostomus Zhang et Yang, 2005
Setihercostomus setifacies (Stackelberg), 1934 – Primorye (Stackelberg, 1934), Khabarovsk territory (Negrobov, Rodionova, 2004 b), Krasnoyarsk Territory (Pogonin, Negrobov, 2008).
Sphyrotarsus Mik, 1874.
Sphyrotarsus caucasicus Negrobov, 1965 – North Caucasus (Negrobov, 1965 b, 1967 c), Adygea (Grichanov, Volfov, Kustov, 2009).
Suschania Negrobov, 2003.
Suschania stackelbergi Negrobov, 2003 – Primorye (Negrobov, 2003).
Sybistroma Meigen, 1824.
Sybistroma binodicornis Stackelberg, 1941 – Leningrad reg. (Stackelberg, 1941), Voronezh reg. (Negrobov, 1977 g), North Caucasus (Grichanov, Volfov, Kustov, 2007), Adygea (Grichanov, Volfov, Kustov, 2009), Sybistroma crinipes Staeger, 1842 – North Caucasus (Grichanov, Volfov, Kustov, 2007), Sybistroma discipes (Germar, 1817) – North Caucasus (Grichanov, Volfov, Kustov, 2007), Sybistroma dufouri Macquart, 1838 – Adygea (Grichanov, Volfov, Kustov, 2009), Sybistroma impar (Rondani, 1843) – Krasnodar Territory (Grichanov, Volfov, Kustov, 2007), Sybistroma nodicornis Meigen, 1928 – Russia (Parent, 1938), Sybistroma obscurella (Fallén, 1823) – North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Adygea (Grichanov, Volfov, Kustov, 2009), Krasnodar Territory (Grichanov, 2012 c), Sybistroma transcaucasica (Stackelberg, 1941) – North Caucasus (Stackelberg, 1941), Adygea (Grichanov, Volfov, Kustov, 2009), Krasnodar Territory (Grichanov, 2012 c).
Sympycnus Loew, 1857.
Sympycnus aeneicoxa (Meigen, 1824) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Sympycnus brevimanus Loew, 1857 – Yakutia (Grichanov, Bagachanova, 2006), Sympycnus cirrhipes (Haliday, 1851) – North Caucasus (Negrobov, 1965 b, Grichanov, Volfov, Kustov, 2007), Krasnodar Territory (Grichanov, Volfov, Kustov, 2006), Altai (Grichanov, 2007 b), Sympycnus pulicarius (Fallén, 1823) (synonym – Sympycnus annulipes (Meigen, 1824)) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915), Novgorod reg. (Stackelberg, 1919), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1972), Murmansk reg. (Negrobov, 1974), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Vologda reg. (Grichanov, 2006 b), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Sympycnus simplicipes Becker, 1908 – Krasnodar Territory (Negrobov, 1965 b, Grichanov, 2004, Grichanov, Volfov, Kustov, 2006, Grichanov, 2012 c), Krasnodar Territory (Negrobov, Duhanina, 1984), North Caucasus (Grichanov, Volfov, Kustov, 2007), Sympycnus simplicitarsis Becker, 1900 – Krasnoyarsk Territory (Becker, 1915), Yakutia (Negrobov, Chalaya, 1991), Sympycnus spiculatus Gerstacker, 1864 – Yakutia (Grichanov, Bagachanova, 2006), Sympycnus urgaicus Negrobov, 1973 – Sayan Mountains (Grichanov, Bagachanova, 2006).
Syntormon Loew, 1857.
Syntormon aulicus (Meigen, 1824) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Voronezh reg. (Negrobov, 1965 c), Syntormon bicolorellus (Zetterstedt, 1843) – Moscow reg. (Fedtchenko A., 1868), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1972), Vologda reg. (Grichanov, 2006 b), Syntormon denticulatus (Zetterstedt, 1843) (synonym – Syntormon pumilus (Meigen, 1824)) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Murmansk reg. (Frey, 1915), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Kursk reg. (Grichanov, 2007 a), Kabardino-Balkaria (Grichanov, Volfov, Kustov, 2007), Ryazan reg. (Negrobov, Pogonin, 2008), Kaluga reg. (Grichanov, 2010 b), Syntormon filiger Verral, 1912 – Krasnodar Territory (Grichanov, Volfov, Kustov, 2006), Astrakhan reg. (Grichanov, 2011), Rostov reg. (Grichanov, 2012 c), Syntormon flexibilis Becker 1922 (synonym – Syntormon lindneri Negrobov, 1975) – Amur reg., Primorye (Negrobov, 1975 b), Syntormon freymuthae Loew, 1873 – Leningrad reg. (Stackelberg, 1925, 1962), Syntormon fuscipes (von Roser, 1840) (synonym Syntormon spicatus (Loew, 1857) – Russia (Parent, 1938), West Caucasus (Negrobov, 1967 c), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2007), Syntormon latitarsis Negrobov et Shamshev, 1984 – North Caucasus (Negrobov, Shamshev, 1984), Syntormon metathesis (Loew, 1850) – Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1972), North Caucasus (Negrobov, 1967 c), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Saratov reg. (Nechay, Negrobov, Volodchenko, 2008), Syntormon monilis (Walker, 1851) – Leningrad reg. (Stackelberg, 1962), North Caucasus (Negrobov, 1967 c), Syntormon monochaetus Negrobov, 1975 – Primorye , Buryatia (Negrobov, 1975 b), Khabarovsk Territory (Grichanov, 2006 a), Syntormon pallipes (Fabricius, 1794) – Murmansk reg. (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Murmansk reg. (Negrobov, 1974), Krasnodar Territory (Negrobov, Duhanina, 1984, Grichanov, Volfov, Kustov, 2006, Grichanov, 2012 c), Syntormon pennatus Ringdahl, 1920 – Kabardino-Balkaria (Grichanov, Volfov, Kustov, 2007), Syntormon pseudopalmare Negrobov et Shamshev, 1985 – Yakutia (Negrobov, Shamshev, 1985), Syntormon pseudospicatus Strobl 1899 – North Caucasus (Negrobov, Shamshev, 1984), Syntormon pumilus (Meigen, 1824) (synonym – Syntormon rufipes (Meigen, 1824)) – Krasnodar Territory (Negrobov, Duhanina, 1984, Grichanov, Volfov, Kustov, 2006), Ryazan reg. (Negrobov, Pogonin, 1984), Vologda reg. (Grichanov, 2006 b), Astrakhan reg. (Grichanov, 2011), Syntormon punctatus (Zetterstedt, 1843) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Vologda reg. (Grichanov, 2006 b), Syntormon sipliviskii Negrobov, 1975 – Buryatia (Negrobov, 1975 b), Syntormon subinermis asiaticus Negrobov, 1975 (Tadzhikistan)– Kabardino-Balkaria (Grichanov, Volfov, Kustov, 2007), Syntormon submonilis Negrobov, 1975 – North Caucasus (Negrobov, 1975 b, Grichanov, Volfov, Kustov, 2007), Syntormon tarsatus (Fallén, 1823) – Murmansk reg. (Frey, 1915), Leningrad reg. (Stackelberg, 1925, 1962), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Kamchatka (Negrobov, Chalaya, 1991), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Syntormon violovitchi Negrobov, 1975 – Far East (Negrobov, 1975b), Kamchatka (Negrobov, Chalaya, 1991).
Systenus Loew, 1857.
Systenus bipartitus (Loew, 1850) – Leningrad reg. (Stackelberg, 1962), Systenus pallipes (von Roser, 1840) – Russia (Parent, 1938), Leningrad reg. (Stackelberg, 1962), Voronezh reg. (Negrobov, 1972), North Caucasus (Grichanov, Volfov, Kustov, 2006), Systenus sachalinensis Negrobov et Shamshev, 1985 – Yakutia (Negrobov et Shamshev, 1985), Systenus scholtzi (Loew, 1850) – North Caucasus (Negrobov, 1977 b).
Tachytrechus Stannius, 1831.
Tachytrechus ammobates (Haliday 1851) (synonym – Tachytrechus plumipes Fallen, 1823) – Siberia (Frey, 1915), Leningrad reg. (Stackelberg, 1921, 1925, 1962), Russia (Parent, 1938), Tachytrechus consobrinus Walker, 1851 – Leningrad reg. (Stackelberg, 1921, 1925), Russia (Parent, 1938), Tachytrechus fedtshenkoi Stackelberg, 1924 – Uzbekistan (Stackelberg, 1924), Tachytrechus genualis Loew, 1857 – Leningrad reg. (Stackelberg, 1925, 1962, Stackelberg, 1941), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Krasnodar Territory (Negrobov, Duhanina, 1984, Grichanov, 2012 c), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010), Tachytrechus hamatus Loew, 1871 – Russia (Loew, 1871), Moscow reg. (Fedtchenko A., 1892), Leningrad reg. (Stackelberg, 1925, 1962, Stackelberg, 1941), Tachytrechus notatus (Stannius, 1831) – Krasnodar Territory (Negrobov, Duhanina, 1984, Grichanov, 2012c), Yakutia (Grichanov, Bagachanova, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2007), Adygea (Grichanov, Volfov, Kustov, 2009, Negrobov, Korneev, 2010), Tachytrechus ripicola Loew, 1857 – Russia (Parent, 1938), Voronezh reg. (Negrobov, 1963, 1965 c, 1966 а, 1967 a), Mordovia (Negrobov, Gоlubcova, 1983), Ural (Negrobov, Radionova, 2004), Krasnodar Territory (Grichanov, Volfov, Kustov, 2006), Tachytrechus transitorius Becker 1917 – North Caucasus (Negrobov, 1965 b, Grichanov, Volfov, Kustov, 2007), Tachytrechus vinogradovi Stackelberg, 1971 – Chita reg. (Stackelberg, 1971), Sayan Mountains (Negrobov, Selivanova, Maslova, 2010).
Telmaturgus Mik, 1874.
Telmaturgus tumidulus (Raddatz, 1873) – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), North Caucasus (Negrobov, 1967 c).
Teuchophorus Loew, 1857.
Teuchophorus bipilosus Becker, 1908 – Krasnodar Territory (Negrobov, Duhanina, 1984, Negrobov, Grichanov, Shamshev, 1984), Teuchophorus calcaratus (Macquart, 1827) – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), North Caucasus (Negrobov, 1967 c), Leningrad reg., Lipetsk reg., North Caucasus (Negrobov, Grichanov, Shamshev, 1984), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), Adygea (Grichanov, Volfov, Kustov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2007), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Adygea (Grichanov, Volfov, Kustov, 2009), Teuchophorus monocanthus Loew, 1859 – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2006, 2007), Voronezh reg. (Negrobov, 1972), Murmansk reg. (Negrobov, 1974), Krasnodar Territory (Grichanov, Kustov, Volfov, 2006, Grichanov, 2012 c), Teuchophorus nigricosta (von Roser, 1840) (synonym – Teuchophorus signatus (Staeger in Zetterstedt, 1849)) – Leningrad reg. (Stackelberg, 1925, 1962), Russia (Parent, 1938), Leningrad reg., Lipetsk reg. (Negrobov, Grichanov, Shamshev, 1984), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Teuchophorus spinigerellus (Zetterstedt, 1843) – Leningrad reg. (Stackelberg, 1925, 1962), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Leningrad reg., North Caucasus (Negrobov, Grichanov, Shamshev, 1984), Kaluga reg. (Grichanov, 2010 b), Teuchophorus ussurianus Negrobov Grichanov & Shamshev, 1984 – Primorye (Negrobov, Grichanov, Shamshev, 1984).
Thinophilus Schiodte, 1844.
Thinophilus argyropalpis Becker, 1910 – Volgograd reg. (Negrobov, 1971 b), Thinophilus flavipalpis (Zetterstedt, 1843) – Rostov reg., North Caucasus (Negrobov, 1971 b), Krasnodar Territory (Grichanov, Volfov, Kustov, 2006, Grichanov, 2012 c), Astrakhan reg. (Grichanov, 2011), Thinophilus longipilus Negrobov, 1971 – Primorye (Negrobov, 1971 b), Thinophilus pollinosus Loew, 1871 – Russia (Loew, 1871), Thinophilus ruficornis (Haliday, 1838) – Murmansk reg., Orenburg reg., Ural, Omsk reg. (Negrobov, 1971 b), Murmansk reg. (Negrobov, 1974), Yakutia (Grichanov, Bagachanova, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007),
Thrypticus Gerstaecker, 1864.
Thrypticus altaicus Negrobov, 1971 – Altai (Negrobov, Stackelberg, 1971, Selivanova, Negrobov, Maslova, 2009, Grichanov, 2007 b), Krasnoyarsk Territory (Negrobov, Tsurikov, 1986 b, Selivanova, Negrobov, Maslova, 2009), Thrypticus aphroditus Negrobov et Thurikov, 1986 – Krasnoyarsk Territory (Negrobov, Tsurikov, 1986 b, Selivanova, Negrobov, Maslova, 2009), Thrypticus atomus Frey, 1915, Arkhangelsk reg. (Frey, 1918), Russia (Parent, 1938), Leningrad reg. (Stackelberg, 1925, 1962, Selivanova, Negrobov, Maslova, 2009), Arkhangelsk reg. (Negrobov, Stackelnerg, 1971), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Yakutia (Grichanov, Bagachanova, 2006), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Thrypticus bellus Loew, 1869 – Leningrad reg. (Stackelberg, 1925, Selivanova, Negrobov, Maslova, 2009), Voronezh reg. (Negrobov, 1963), Siberia (Negrobov, Stackelberg, 1971), Krasnodar Territory (Grichanov, Volfov, Kustov, 2006), North Caucasus (Grichanov, Volfov, Kustov, 2007), Krasnoyarsk Territory (Pogonin, Negrobov, 2008), Thrypticus cuneatus (Becker, 1917) – Russia (Becker, 1917), Leningrad reg. (Stackelberg, 1925, 1962, Selivanova, Negrobov, Maslova, 2009), Thrypticus divisus (Strobl, 1880) – Leningrad reg. (Selivanova, Negrobov, Maslova, 2009), Thrypticus emiliae Negrobov, 1972 – Altai (Negrobov, Stackelberg, 1972, Selivanova, Negrobov, Maslova, 2009, Grichanov, 2007 b), Yakutia (Grichanov, Bagachanova, 2006), Thrypticus incanus Negrobov, 1967 – Leningrad reg. (Negrobov, 1967 b, Negrobov, Stackelberg, 1972, Selivanova, Negrobov, Maslova, 2009), Thrypticus intercedens Negrobov, 1967 – Yaroslavl reg. (Negrobov, 1967 b), Sayan Mountains (Negrobov, Stackelberg, 1972, Selivanova, Negrobov, Maslova, 2009), Yakutia (Grichanov, Bagachanova, 2006), Yakutia, Khanty-Mansi autonomous region (Grichanov, 2010 a), Thrypticus laetus Verrall, 1912
– Leningrad reg. (Stackelberg, 1962, Negrobov, Stackelberg, 1972), Voronezh reg. (Negrobov, 1972, Selivanova, Negrobov, Maslova, 2009), Thrypticus nigricauda Wood, 1913 – Leningrad reg. (Stackelberg, 1925, 1962, Selivanova, Negrobov, Maslova, 2009), Voronezh reg. (Negrobov, 1963, Negrobov, Stackelnerg, 1972), Thrypticus paludicola Negrobov, 1972 – Sayan Mountains (Negrobov, 1972), Thrypticus politus Negrobov, 1967 – Leningrad reg. (Negrobov, Stackelberg, 1972), Thrypticus pollinosus Verrall, 1912 – Leningrad reg. (Stackelberg, 1925, 1962, Selivanova, Negrobov, Maslova, 2009), Voronezh reg. (Negrobov, 1963), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Thrypticus riparius Negrobov, 1972 – Primorye (Negrobov, Stackelberg, 1972, Selivanova, Negrobov, Maslova, 2009), Thrypticus smaragdinus Gerstaecker, 1864 – Krasnodar Territory (Selivanova, Negrobov, Maslova, 2009), North Caucasus (Grichanov, Volfov, Kustov, 2006, 2007), Thrypticus tarsalis Parent, 1932 – Leningrad reg. (Stackelberg, 1962, Negrobov, Stackelberg, 1972, Selivanova, Negrobov, Maslova, 2009), Thrypticus tonsus Negrobov, 1972 – (Karaganda) (Negrobov, Stackelberg, 1972), Krasnodar Territory (Selivanova, Negrobov, Maslova, 2009), Thrypticus tsacasi Negrobov, 1972 – Amur reg. (Negrobov, Stackelberg, 1972, Selivanova, Negrobov, Maslova, 2009), Thrypticus virescens Negrobov, 1967 – Leningrad reg. (Negrobov, 1967 b, Negrobov, Stackelberg, 1972, Selivanova, Negrobov, Maslova, 2009), North Caucasus (Grichanov, Volfov, Kustov, 2006).
Vetimicrotes Dyte, 1980.
Vetimicrotes baskunchakensis Grichanov, 2011 – Astrakhan reg. (Grichanov, 2011).
Xanthochlorus Loew, 1857.
Xanthochlorus fulvus Negrobov, 1978 – North Caucasus (Negrobov, 1978 a), Adygea (Grichanov, Volfov, Kustov, 2006, 2009), Krasnodar Territory (Grichanov, 2012 c), Xanthochlorus luridus Negrobov, 1978 – North Caucasus (Negrobov, 1978 a), Adygea (Grichanov, Volfov, Kustov, 2009), Xanthochlorus philippovi Negrobov, 1978
– Far East (Negrobov, 1978 a), Xanthochlorus tenellus (Wiedemann, 1817) – Moscow reg. (Fedtchenko A., 1868, Fedtchenko B., 1892), Leningrad reg. (Stackelberg, 1925, 1962), Voronezh reg. (Negrobov, 1963, 1965 c, 1967 a), North Caucasus (Negrobov, 1967 c, Grichanov, Volfov, Kustov, 2007), Murmansk reg. (Negrobov, 1974), Ryazan reg. (Negrobov, Pogonin, 1984, Negrobov, Pogonin, 2008), Adygea (Grichanov, Kustov, Volfov, 2006, 2009), Tatarstan (Negrobov, Korneev, Selivanova, 2010).
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Negrobov, O.P., 1963. K izucheniju naibolee poleznyh entomofagov iz semejstva muh dolihopodid Voronezhskoj oblasti // Sbornik nauchnyh studencheskih rabot. Voronezh, vyp. 2. S. 1-3. Negrobov, O.P., 1965 a. K izucheniju endemichnyh vidov Dolichopodidae (Diptera) Kavkaza // Sbornik nauchnyh rabot aspirantov VGU. Voronezh, vyp. 2. S. 118-121. Negrobov, O.P., 1965 b. Novye i maloizvestnye vidy Dolichopodidae (Diptera) fauny SSSR // Entomol. obozr., № 44, vyp. 2. S. 438-446. Negrobov, O.P., 1965 c. K izucheniju semejstva Dolichopodidae v pojmennyh cenozah Voronezhskoj oblasti // Biologicheskie osnovy rekonstrukcii. Kishinjov. S. 409-411. Negrobov, O.P., 1966 a. K izucheniju Dolichopodidae (Diptera) Voronezhskoj oblasti // Referaty dokladov IV nauchnoj konferencii molodyh uchjonyh-biologov. Moskva. S. 75-76. Negrobov, O.P., 1966 b. Revizija palearkticheskih vidov roda Medetera (Dolichopodidae, Diptera). Soobshhenie I. Gruppa Oligochaetus Mik// Entomol. Obozr., №45, vyp. 4. S. 874-896. Negrobov, O.P., 1967 a. K faune Dolichopodidae (Diptera) Usmanskogo bora // Zoologicheskij institut AN SSSR, vyp. 15. S. 79-82. Negrobov, O.P., 1967 b. Novye palearkticheskie vidy podsemejstva Medeterinae (Dolichopodidae, Diptera) // Entomologicheskoe obozrenie, T. XLVI, vyp. 4. S. 890-908. Negrobov, O.P., 1967 c. Fauna i vertikal'noe raspredelenie Dolichopodidae (Diptera) Zapadnogo Kavkaza // Zool. zhurnal, №46, vyp. 7. S. 1047-1079. Negrobov, O.P., 1968 a. Dva novyh vida hishhnyj dvukrylyh roda Medetera Fischer (Diptera, Dolichopodidae) // Doklady akademii nauk Armjanskoj SSR. T. XLV. S. 188-192. Negrobov, O.P., 1968 b. K izucheniju uglevodorodnogo pitanija semejstva Dolichopodidae (Diptera)// Sbornik rabot aspirantov VGU. Voronezh, vyp. 4. S. 240-243. Negrobov, O.P., 1968 c. Novyj rod i vidy Dolichopodidae (Diptera) // Zoologicheskij zhurnal, T. 43, vyp. 3. S. 470-472. Negrobov, O.P., 1970. A contribution to the knowledge of Medetera of Japan (Diptera, Dolichopodidae)// Insecta Matsumurana, №9. P. 1-7. Negrobov, O.P., 1970 b. Sluchai gornogo melanizma u vidov semejstva Dolichopodidae (Diptera) Zapadnogo Kavkaza // Trudy VGU. Voprosy zoologii, fiziologii i biofiziki. Voronezh, t.
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Negrobov, O.P., 1973 b. Novye vidy roda Dolichopus (Diptera, Dolichopodidae) Vostochnogo Sajana, Pribajkal'ja i Zabajkal'ja // Novye i maloizvestnye vidy fauny Sibiri. Novorossijsk, vyp. 7. S. 129-133.
Negrobov O.P., 1973 c. Novye vidy dvukrylyh roda Argyra (Diptera, Dolichopodidae) fauny SSR // Dvukrylye dal'nego vostoka, №5. S. 5-9.
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Negrobov, O.P., 1974. K faune Dolichopodidae, Diptera Laplandskogo i Kandalakshskogo zapovednika // Problemy izuchenija i ohrany landshaftov. Voronezh. S. 43-46.
Negrobov, O.P., 1975 a. Eine neue Art der Gattung Hydrophorus Fall. (Diptera, Dolichopodidae) aus der Sammlung des Zoologischen Museums Helsinki. P. 10-12.
Negrobov, O.P., 1975 b. Obzor muh-zelenushek roda Syntormon Meigen (Diptera, Dolichopodidae) fauny Palearktiki // Entomologicheskij obzor, T. LIV, vyp. 3. S. 652-664.
Negrobov, O.P., 1975 c. Vidy roda Hydrophorus (Dolichopodidae, Diptera) fauny SSSR // Problemy izuchenija i ohrany landshaftov. Voronezh. Vyp. 2. S. 63-66.
Negrobov, O.P., 1976 a. Novye vidy roda Rhaphium (Dolichopodidae, Diptera) Sibiri i Dal'nego Vostoka // Zoologicheskij zhurnal, T. LV, vyp. 6. S. 860-865.
Negrobov, O.P., 1976 b. Novye i maloizvestnye vidy semejstva Dolichopodidae (Diptera) fauny SSSR i sopredel'nyh territorij // Nauchnye doklady vysshej shkoly, biologicheskie nauki, №8. S. 45-50.
Negrobov, O.P., 1976 c. Vidy Dolichopus podroda Hydroceleuthus (Diptera, Dolichopodidae) vostochnoj Palearktiki // Nasekomye Mongolii. L. Vyp. 4. S. 509-517.
Negrobov, O.P., 1977 a. Dolichopodidae. In E. Lindner // Die Fliegen der palearctiscen Region. Bd. IV, 5, 29, Lief. 316. P. 347-386.
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Negrobov, O.P., 1977 c. Novye vidy roda Dolichopus Lart. (Diptera, Dolichopodidae) Sibiri i Dal'nego Vostoka // Taksony fauny Sibiri i Dal'nego Vostoka. – Novosibirsk. S. 92-104.
Negrobov, O.P., 1977 d. Novye vidy roda Rhaphium Mg. (Dolichopodidae, Dipterta) fauny SSSR // Vestnik zoologii, №1. S. 45-49.
Negrobov, O.P., 1977 e. Novyj rod semejstva Dolichopodidae (Diptera) iz Primor'ja. Entomofauna Dal'nego Vostoka // Trudy Biologo-pochvennogo instituta, T. 46, vyp. 149. S. 146
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Negrobov, O.P., 1977 g. Uber die Fauna der Familie Dolichopodidae (Diptera) des Zentralen Schwarzerdegebiets // VII Mezhdunarodnyj simpozium po entomofaune Srednej Evropy. Leningrad. S. 71.
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Negrobov, O.P, 1986 a. K faune semejstva Dolichopodidae (Diptera) Amurskoj oblasti // Trudy zoologicheskogo institute AN SSSR, T. 146. S. 10-14. Negrobov, O.P., 1986 b. K faune semejstva Dolichopodidae (Diptera) Amurskoj oblasti SSSR i Mongolii // Trudy zoologicheskogo instituta AN SSSR. T. 146. S. 35-38. Negrobov, O.P., 1987. Novye palearkticheskie vidy dolihopodid rody Neurigona Rond. (Diptera, Dolichopodidae) // Entomologicheskoe obozrenie, T. LXVI, vyp. 2. S. 406-415.
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Negrobov, O.P., Barkalov, A.V., 1978. Novye vidy roda Dolichopus Latr. (Dolichopodidae, Diptera) Sibiri, Primor'ja i Yakutiaa // Taksonomija i ekologija chlenistonogih Sibiri. Novosibirsk. S. 154-162.
Negrobov, O.P., Barkalov, A.V., 2009. Novye dannye po faune i sistematike vidov semejstva Dolichopodidae (Diptera) gor Altaja s opisaniem novogo vida // Altaiskij zoologicheskij zhurnal, vyp. 3. S. 3-12.
Negrobov, O.P., Barkalov, A.V., 2010. Novye vidy roda Dolichopus (Diptera, Dolichopodidae) s gor Altaja i Tuvy // Zoologicheskij zhurnal, T. 89, №11. S. 1361-1365.
Negrobov, O.P., Bulli, A.F, 1986. Maloizvestnye vidy semejstva Dolichopodidae (Diptera) iz zaporozhskoj oblasti // Voronezh. gos. un-t. S. 5.
Negrobov O.P., Chalaja, O.N., 1987. Novye dannye o rode Hercostomus Loew (Dolichopodidae, Diptera) SSSR // Nauchnye doklady vysshej shkoly, Biol. nauki. № 5. S. 42-46.
Negrobov, O.P., Chalaja O.N., 1988. Palearkticheskie vidy roda Lamprochromus (Diptera, Dolichopodidae) // Zoologicheskij zhurnal, T.67, vyp. 5. S. 795-799.
Negrobov, O.P., Chalaja O.N., 1991. Dolihopodidy (Dolichopodidae, Diptera) Severo-Vostochnoj chasti SSSR // Entomologicheskoe issledovanija na severo-vostoke SSSR. Voronezh.
S. 66-80. Negrobov, O.P., Fursov, V.N., 1988. Revizija vidov roda Neurigona Rond. (Diptera, Dolichopodidae) palearktiki. II // Entomologicheskoe obozrenie, T. LXVII, vyp. 2. S. 405-416. Negrobov, O.P., Golubcov, D.N., 1993. Novye vidy roda Medetera Fischer (Dolichopodidae, Diptera) // Voronezh. gos. un-t. S. 50-53. Negrobov, O.P., Golubtzov, D.N., 2006. A new species of the genus Hydrophorus (Diptera, Dolichopodidae) from the Far East of Russia // Studia dipterologica 12 (2). P. 473-475.
Negrobov, O.P., Golubcova, T.N, 1983. Nekotorye dannye po faune vidov semejstva Dolichopodidae (Diptera) Mordovii // Voprosy fauny i ekologii nasekomyh, Voronezh, 18-20 marta 1980: Materialy nauch. konf. Voronezh. gos. un-t. S. 25-28.
Negrobov, O.P., Grichanov I.Ya., Barkalov, A.V., 2009. The Dolichopus latipennis species group (=Hygroceleuthus Loew) in the Palearctic Region (Diptera: Dolichopodidae) // Zootaxa 2087. P. 37-45.
Negrobov, O.P, Grichanov, I.Ya., Shamshev, I.V., 1984. Obzor palearkticheskih vidov roda Teuchophorus Loew (Dolichopodidae, Diptera) // Nauch. dokl. vyssh. shk. Biol. nauki, № 8. S. 37-42.
Negrobov, O.P., Duhanina, E.V., 1984. K izucheniju vidov semejstva Dolichopodidae (Diptera) okrestnostej Gelendzhika // Voronezh. gos. un-t. S. 1-5.
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Negrobov, O.P., Korneev I.I., 2010. Novye dannye po faune vidov semejstva Dolichopodidae (Diptera) Evropejskoj chasti Rossii // «Aktual'nye voprosy sovremennoj entomologii i ekologii nasekomyh»: Materialy mezhdunarodnoj nauchnoj konferencii, posvjashhjonnoj pamjati A.I. Fomichjova, Borisoglebsk, 3-4 dekabrja 2009 g. Borisoglebsk. S. 79-80
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Negrobov, O.P., Maslova, O.O., 2003. Novye vidy roda Argyra (Diptera, Dolichopodidae) Rossii i Kazahstana // Zoologicheskij zhurnal, T. 82, №12. S. 1522-1524.
Negrobov, O.P., Maslova, O.O., 2005. Novyj vid roda Diaphorus (Dolichopodidae, Diptera) s Severnogo Kavkaza // Bjulleten' Moskovskogo obshhestva ispytatelej prirody, Otdelenie Biologii, t.110, vyp.3. S. 77-78.
Negrobov, O.P., Maslova, O.O., 2005. Re-description of a some species from the genus Diaphorus (Dolichopodidae, Diptera) described by A.A. Stackelberg // Int. J. Dipterol. Res. 16(3).
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Negrobov, O.P., Maslova, O.O., Selivanova, O.V., 2005. Redescriptions of the types of species from the genus Diaphorus described by Loew and Becker from the Palaearctic Region (Dolichopodidae, Diptera) // Int. J. Dipterol. Res. 17(1). P. 57-63.
Negrobov, O.P., Maslova, O.O., Selivanova, O.V., 2010. Fauna of the family Dolichopodidae (Diptera) of Sayan Mountains // Int. J. Dipterol. Res. 21(3). P. 175-178.
Negrobov, O.P., Maslova, O.O., Selivanova, O.V., 2011. New and little known species of the genus Dolichopus Latreille, 1796 (Diptera, Dolichopodidae) from Russia // Far Easten Entomologist, N 232. P. 11-16.
Negrobov, O.P., Nechaj, N.A., 2009. Novyj vid roda Hercostomus Loew, 1857 (Diptera, Dolichopodidae) s opredelitel'noj tablicej palearkticheskih vidov gruppy IV // Bjulleten' Moskovskogo obshhstva ispytatelej prirody. Otd. biologii, t. 114, vyp. 5. S. 84-87.
Negrobov, O.P., Nechaj, N.A., 2011. Novye dannye po sistematike grupp vidov Hercostomus nigriplantis (Stannius, 1831) (Diptera, Dolichopodidae) // Evrazijskij entomologicheskoj zhurnal, № 10, vyp. 1. S. 121-123.
Negrobov, O.P., Pogonin, S.V., 1984. K izucheniju vidov semejstva Dolichopodidae (Diptera) Okskogo zapovednika // Voronezh. gos. un-t. S. 1-10.
Negrobov, O.P., Pogonin, S.V., 2006. Fauna vidov roda Rhaphium Meiger, 1803 (Diptera, Dolichopodidae) Krasnojarskogo kraja // Kavkazskij entomologicheskij bjulleten', №2, vyp. 2. S. 219-220.
Negrobov, O.P., Pogonin, S.V., 2008. On the species of the family Dolichopodidae (Diptera) from the Ryazan region // Int. J. Dipterol. Res. 19(2). P. 97-100.
Negrobov, O.P., Rodionova, S.Y., 2004 а. New data on fauna of subfamily Dolichopodinae (Dolichopodidae, Diptera) in Russia and neighbouring territories (genera Dolichopus Latr. and Tachytrechus Walk.) // Int. J. Dipterol. Res. 15(3). P. 191-196.
Negrobov, O.P., Rodionova, S.Y., 2004 b. New data on fauna of subfamily Dolichopodinae (Dolichopodidae, Diptera) in Russia and neighbouring territories (genus Hercostomus Lw.) // Int.
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Negrobov, O.P., Sviridova E.L., 1983. K izucheniju fauny muh roda Dolichopus Latr. (Dolichopodidae, Diptera) Primor'ja // Sistematika i ekologo-faunisticheskij obzor otdel'nyh otrjadov nasekomyh Dal'nego Vostoka. S. 110-114.
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Negrobov, O.P., Selivanova, O.V., 2005. Novyj vid roda Argyra Macquart (Diptera, Dolichopodidae) s Severnogo Kavkaza // Bjulleten' Moskovskogo obshhestva ispytatelej prirody. Otd. biol. T. 110, vyp. 3. S. 70-71.
Negrobov, O.P., Selivanova, O.V., 2010. A new species of the genus Achalcus Low, 1857 (Diptera: Dolichopodidae) from Far East region of Russia // Кавказский энтомол. бюллетень №6, вып. 1. S. 123−124.
Negrobov, O.P., Shamshev, I.V., 1982. Novye dannye po semejstvu Dolichopodidae (Diptera) juga evropejskoj chasti SSSR // Voprosy fauny i ekologii nasekomyh, Voronezh, 18-20 marta 1980: Materialy nauchnoj konferencii VGU. S. 19-24.
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Работа выполнена при поддержкегранта РФФИ (11-04-01051 а).
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Оглавление
Contents
Гричанов И.Я. Заметки по систематике западно-палеарктических видов рода Syntormon Loew (Diptera: Dolichopodidae) ........................3 Grichanov I.Ya. Systematic notes on West-Palaearctic species of the genus Syntormon Loew (Diptera: Dolichopodidae) .....................................3 Гричанов И.Я. Видырода Asyndetus Loew (Diptera: Dolichopodidae) тропической Африки с заметками о некоторых палеарктических и ориентальных видах ..............................................................................27 Grichanov I.Ya. Afrotropical species of the genus Asyndetus Loew (Diptera: Dolichopodidae) with notes on some Palaearctic and Oriental species ..........................................................................................................27 НегробовО.П., Селиванова О.В., Маслова О.О., Чурсина М.А. Справочный список хищных мух семейства Dolichopodidae (Diptera) фауны России ........................................................................................47 Negrobov, O.P., Selivanova, O.V., Maslova, O.O., Chursina M.A. Check-list of predatory flies of the family Dolichopodidae (Diptera) in the fauna of Russia .................................................................................47
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В.Г.Иващенко, Н.П.Шипилова, Л.А.Назаровская. Фузариоз колоса хлебных злаков. СПб, 2004.
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Igor Ya. Grichanov. A checklist and keys to North European genera and species of Dolichopodidae (Diptera). St.Petersburg, 2006.
Igor Ya. Grichanov. A checklist and keys to Dolichopodidae (Diptera) of the Caucasus and East Mediterranean. St.Petersburg, 2007.
Лаборатория микологии и фитопатологии им. А.А.Ячевского ВИЗР. История и современность. Под редакцией А.П.Дмитриева. СПб, 2007.
В.В. Нейморовец. Полужесткокрылые насекомые (Heteroptera) Краснодарского края и Республики Адыгея. Список видов. Под редакцией И.Я.Гричанова. СПб, 2010.
Фауна и таксономия хищных мух Dolichopodidae (Diptera). Сборник научных работ. Подредакцией И.Я.Гричанова иО.П.Негробова. СПб, 2013.
Научноеиздание. RIZO-печать ООО "ИННОВАЦИОННЫЙ ЦЕНТР ЗАЩИТЫРАСТЕНИЙ" Лицензия ПЛД №69-253. Подписанокпечати 11 марта 2013 г.
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